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Mandarin duck (Aix galericulata) drake standing on frozen pond during snowstorm. Richmond Park, London, UK. January
Male Black woodpecker (Dryocopus martius) in snow, showing red crown. Kuusamo Finland December. Overall winner of the Swiss Vogelwarte photo competition 2021
Greater Prairie-Chicken (Tympanuchus cupido) male displaying on lek. Fort Pierre National Grassland, South Dakota, USA. When displaying
Common kingfisher (Alcedo atthis) perched on a bulrush, Devpm, UK, November
Great horned owls (Bubo virginianus), adult and chick, on nest in Saguaro cactus (Carnegiea gigantea), Sonoran desert, Arizona
Tawny owls (Strix aluco) peering out from tree hollow, Dumfries and Galloway, Scotland, UK, May, 2021
Tawny owl (Strix aluco) perched in Silver Birch tree (Betula pendula), Dumfries and Galloway, Scotland, UK, May
Superb lyrebird (Menura novaehollandiae) male in partial display perched in Tree fern (Cyatheales) in wet eucalypt forest, Lakes Entrance Rainforest, Victoria, Australia
Crested tit (Lophophanes cristatus), Sierra de Grazalema Natural Park, Southern Spain. September
Mandarin duck (Aix galericulata) male swimming with autumn colours reflected in the water, London, UK, November
Song thrush (Turdus philomelos) looking for worms in a garden, Broxwater, Cornwall, UK. April
Great horned owl (Bubo virginianus) adult and chick in nest in Saguaro cactus (Carnegiea gigantea) Sonoran Desert, Arizona, USA
Bearded vulture (Gypaetus barbatus) flying, Ordesa National Park, Pyrenees, Spain
Two Long-eared owl chick (Asio otus) perched in tree. Alsace. France
Baya weaver (Ploceus philippinus) breeding male perched on nest, rural Karantaka, India
An adult King penguin (Aptenodytes patagonicus) amongst a creche of chicks at the breeding colony. Fortuna Bay, South Georgia Island
A breeding pair of King penguins (Aptenodytes patagonicus) with their chick. St Andrews Bay, South Georgia Island
King penguins (Aptenodytes patagonicus) congregate on the beach, seeking a safe point to go into the sea, with male Southern elephant seals (Mirounga leonina) blocking their path
Woodcock (Scolopax rusticola), in undergrowth in winter, Brittany, France
Red-footed booby (Sula sula), brown morph, Genovesa Island, Galapagos Islands
Galapagos penguin (Spheniscus mendiculus), pair courting, Isabela Island, Galapagos Islands
Galapagos hawk (Buteo galapagoensis), juvenile plumage, Bainbridge Crater, Galapagos Islands
Darwins cactus finch (Geospiza scandens), feeding on Opuntia cactus flower, Santa Cruz Island, Galapagos Islands
Kingfisher ( Alcedo atthis) male and female perched on a root when building the nest, Lorraine, France, March
Pied avocet (Recurvirostra avosetta) adult parent and chick together. Suffolk, UK. June
Mistle thrush (Turdus viscivorus) calling perched on rowan tree. London, UK. December
Satin bowerbird (Ptilonorhynchus violaceus), maintaining his bower, Ravenshow, North Queensland, Australia, October
Eastern rockhopper penguins (Eudyptes chrysocome filholi), Auckland Island, December
Crested tit (Lophophanes cristatus) silhouette, Sierra de Grazalema Natural Park, Cadiz, Andalusia, Southern Spain. October
Rowi or Okarito Brown Kiwi (Apteryx rowi) male called Scooter in rainforest habitat. Okarito Forest, Westland, South Island, New Zealand
Mute swan (Cygnus olor) in mist, Bolam Lake Country Park, Northumberland, UK, November
Aerial view of green algae and Lesser flamingos (Phoeniconaias minor) in flight at the mouth of a river, Lake Bogoria, Kenya
Common swift (Apus apus) flying low to water while hunting, Norfolk, England, UK, July. Winner of Animal Behaviour Category of the British Wildlife Photography Awards 2019. Photographer
White-tailed sea eagle (Haliaeetus albicilla) flying over water with male Common eider prey (Somateria mollissima), Uto, Parainen, Finland, April
Male Black grouse (Tetrao / Lyrurus tetrix) in flight. Kuusamo, Finland, April
Laysan albatross (Phoebastria immutabilis) chick in the nest, Guadalupe Island Biosphere Reserve, off the coast of Baja California, Mexico, February
Buzzard (Buteo buteo) landing on branch, winter, Lorraine, France, February
Boat-tailed grackle pair (Quiscalus major) in courtship display. Wakodahatchee Wetlands, Florida, USA, April
Aerial view of Griffon vulture (Gyps fulvus) flock scavenging dead pig, Cantabria, Spain. January
Ruby-throated hummingbird (Archilochus colubris) male landing on Virginia creeper (Parthenocissus quinquefolia). Hill Country, Texas, USA
Kingfisher (Alcedo atthis) male, after diving, taking off from water with fish, a Common Roach (Rutilus rutilus) Lorraine, France, July
White-tailed eagle (Haliaeetus albicilla) flying over water with fish in talons. Norway. August
Great grey owl (Strix nebulosa) in flight in woodland during snowfall. Kuusamo. Finland. February
Barred Owl (Strix varia) fledglings. Washington County, Oregon, USA. June
Griffon vulture (Gyps fulvus) soaring over Los Mallos de Riglos, Huesca province, Aragon, Spain
African fish eagle (Haliaeetus vocifer) swoops to catch a freshly caught fish, dropped by a Saddle-billed stork (Ephippiorhynchus senegalensis) after being pressurised to do so by the eagle
Taiwan Blue Magpie, (Urocissa caerulea) endemic species, Taipei city park, Taiwan
Malayan night heron (Gorsachius melanolophus) Taipei city park, Taiwan
