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Red-footed booby (Sula sula), pair perched in tree. Genovesa Island, Galapagos
Red-footed booby (Sula sula), pair nest building in tree. Darwin Bay, Genovesa Island, Galapagos
Nazca booby (Sula granti) at nest with egg. Overlooking sea, Gardner Islet, Floreana Island, Galapagos. December 2014
Nazca booby (Sula granti), pair amongst Prickly pear (Opuntia sp) cacti at coast. Wolf Island, Galapagos. August 2016
Nazca booby (Sula granti), pair billing. Genovesa Island, Galapagos. May 2015
Lapwing (Vanellus vanellus) portrait in upland hay meadow, Upper Teesdale, CO Durham, England, UK, June
Toco toucan (Ramphastos toco) acrobatically perched, feeding in forest canopy. Northern Pantanal, Cuiaba River, Mato Grosso, Brazil
Blue-footed booby (Sula nebouxii), six diving into sea. Espumilla Beach, Santiago Island, Galapagos
Trumpeter swans (Cygnus buccinator) on the edge of the Upper Yellowstone River. Hayden Valley, Yellowstone National Park, Wyoming, USA. January
Burrowing owl (Athene cunicularia) doing rain dance, spreading wings and dancing while it rained, Arizona, USA
Cuckoo (Cuculus canorus) male, Germany, April
Blackcap (Sylviya atricapilla) male perched in blossom, Hungary, April
Red-collared lorikeet (Trichoglossus rubritorquis) pair examining hole in Eucalyptus as a potential nesting hollow. Katherine, Northern Territory, Australia
Great bowerbird (Chlamydera nuchalis), male showing off treasures whilst female inspects bower. Lake Argyle, Kununurra, Western Australia
Burrowing owl (Athene cunicularia) rotating head, Arizona, USA
Kestrel (Falco tinnunculus) fighting Hungary, April
Common Snipe (Gallinago gallinago) autumn courtship display, Germany, August
Blackbird (Turdus merula) in flight to feed on berries, Helsinki, Finland. November
Bearded reedling (Panurus biarmicus) on frosty reed seed head, Helsinki, Finland, January
Great grey owl (Strix nebulosa) taking off from snowy ground, Kuusamo, Finland. March
Red-legged honeycreeper (Cyanerpes cyaneus) two males on barbed wire. Costa Rica
Flock of Tree sparrows (Passer montanus) in flight, with a Greenfinch (Chloris chloris) blurred motion, Pacsmag, Hungary
Atlantic puffin (Fratercula arctica) portrait with beak open / calling. Shetland IslandsAtlantic puffin (Fratercula arctica) portrait with beak open/calling. Shetland Islands, Scotland, July
Male Golden Pheasant (Chrysolophus pictus) standing on grass in display plumage. Kew Gardens, England, UK. March
Pine grosbeak (Pinicola enucleator), male feeding juvenile, Finland, July
Black grouse (Lyrurus tetrix), male at lek, Finland, April
Long-tailed duck (Clangula hyemalis), males in flight, Finland, April
Kingfisher (Alcedo atthis) male, after diving, taking off from water with fish, a Common Roach (Rutilus rutilus) Lorraine, France, August
Common swift (Apus apus) group flying overhead, Bradford-on-Avon, Wiltshire, UK, May
White-throated magpie-jay (Calocitta formosa) Nicoya peninsula, Costa Rica. February
Pairs of Scarlet Macaws (Ara macao) in flight. Osa Peninsula (near Corcovado National Park), Costa Rica, Central America
Eurasian hoopoe (Upupa epops) at nest in stone wall with hungry chicks gaping, Vendee, France, June
Scarlet ibis (Eudocimus ruber), two birds perched on bushes, Coro, Venezuela. Small repro only
Scarlet ibis (Eudocimus ruber), in flight, Coro, Venezuela
Emerald toucanet (Aulacorhynchus prasinus) Rio Platano Biosphere Reserve and UNESCO World Heritage Site, La Mosquitia, Honduras
Whale headed / Shoebill (Balaeniceps rex) vocalising. Swamps of Mabamba, Lake Victoria, Uganda
Stellers Sea Eagle (Haliaeetus pelagicus) with prey in beak in mid-air, Hokkaido, Japan, February
Harpy eagle (Harpia harpyja) close up head portrait, captive
Lammergeier / Bearded vulture (Gypaetus barbatus) flying, Leukerbad, Valais, Switzerland, February. Highly commended in the Portfolio category of the Terre Sauvage Nature Images Awards 2017
Song Thrush (Turdus philomelos), Bayern, Germany. June. June
Dalmatian pelicans (Pelecanus crispus) underwater view of group feeding, Lake Kirkini, Greece. Highly commended in the GDT competition 2013
King penguins (Aptenodytes patagonicus) at sunrise. Two males and a female, with the males fighting for the attention of the female. Falklands Islands
Andean cock-of-the-rock (Rupicola peruvianus), two males perched in tree at lek in mid-altitude montane rainforest. Manu Biosphere Reserve, Amazonia, Peru
Snowy egret (Egretta thula) portrait, viewed from below. St. Petersburg, Florida, USA
RF - Andean cock-of-the-rock (Rupicola peruvianus), male perched in tree at a lek. Mid-altitude montane rainforest, Manu Biosphere Reserve, Amazonia, Peru
Red-and-green macaw (Ara chloropterus) flock feeding on wall of clay lick. Manu Biosphere Reserve, Amazonia, Peru
Blackbird (Turdus merula), male singing whilst perched in Ash (Fraxinus excelsior) tree, near Bath, England, UK. May
European roller (Coracias garrulus) peering out of nest with feather, Lake Csaj, Pusztaszer, Kiskunsagi National Park, Hungary, February