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Aerial of Kossol Passage, Kossol Reef, Palau Islands, North Pacific Ocean
Palmar tree frog (Boana / Hypsiboas pellucens) on plant stem, Mindo, Pichincha, Ecuador
Andean forest pit viper (Bothriopsis pulchra) curled up on fern, Sumaco, Napo, Ecuador
Booted racket tail (Ocreatus underwoodii) two males fighting in flight, Mindo, Pichincha, Ecuador
Eyelash palm pit viper (Bothriechis schlegelii) with tongue extended, Mindo, Pichincha, Ecuador
Parsons Chameleon (Calumma parsonii) close up of eye and skin, Andasibe-Mantadia National Park, Moramanga, Madagascar
Magnificent land-snail (Helicophanta magnifica) Andasibe-Mantadia National Park, Moramanga, Madagascar
Spotted fish-scaled gecko (Geckolepis maculata) close up of scales, Palmarium, Atsinanana, Madagascar
Bigfin squid (Sepioteuthis lessoniana) hovering in mid water at night. Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
Yellow lipped sea snake (Laticauda colubrina) dives back down to the reef after surfacing for breath of air. Apo Island, Dumaguete, Negros, Philippines. Bohol Sea, Tropical West Pacific Ocean
RF- Pair of Rock beauties (Holacanthus tricolor) spawning at dusk. The larger male rubs against the flank of female during the spawning rise. Cayman Islands, British West Indies. Caribbean Sea
Secretary blenny (Acanthemblemari maria) yawns as it peers out from a hole in the reef. West Bay, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
Boulder brain coral (Colpophyllia natans) photographed with a long exposure with camera rotation. Taken with circular fisheye lens. Grand Cayman, Cayman Islands. Caribbean Sea
Aggregation of tiny isopods (Santia sp. ) living on the surface of a Blue sponge (Haliclona sp)Aggregation of tiny isopods (Santia sp.) living on the surface of a Blue sponge (Haliclona sp.), there are two colour varieties here, possibly different species
Southern stingrays (Hypanus americanus) swimming over a sand bar in the early morning. Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea
Lionfish (Pterois volitans) portrait, invasive species in the Caribbean. East End, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea
Close up of pattern of the cerata of a nudibranch (Janolus savinkini). Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
Hairy frogfish (Antennarius striatus) on a sandy seabed. Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
Nassau grouper (Epinephelus striatus) yawning on a coral reef drop off. San Salvador island, Bahamas. Bahamas Sea, Tropical West Atlantic Ocean
A stand of Pillar coral (Dendrogyra cylindrus) growing on a coral reef, East End, Grand Cayman, Cayman Islands
Bigfin reef squid (Sepioteuthis lessoniana) capturing a pelagic shrimp with long red antennae, in midwater, at night. Anilao, Batangas, Luzon, Philippines
Orange-finned anemonefish (Amphiprion chrysopterus) guarding their recently spawned red eggs on a rock next to the host anemone. Rock Islands, Palau, Micronesia. Tropical west Pacific Ocean
Coleman shrimp (Periclimenes colemani) close up, on a fire urchin (Asthenosoma varium). Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
Yellowhead jawfish (Opistognathus aurifrons) spitting out sand as it digs its burrow. East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
Parachute daisies (Ursinia anthemoides) Little Karoo, Western Cape, South Africa
Yellow tube sponge (Aplysina fistularis) on a coral reef, with the camera rotated during the long exposure. East End, Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea
Kelp gull (Larus dominicanus) on rocky shore with crashing waves at sunset, Bird Island, Lamberts Bay, Western Cape province, South Africa, September 2012
Tropical forest, Andasibe-Mantadia National Park, Madagascar, March 2005
Detail of palm leaf, backlit in sunshine, New Caledonia
RF- Boulder star coral (Montastrea annularis) showing fluorescent green coloration, photographed under blue light at night, to excite fluorescent pigments in the coral, East End, Grand Cayman
Southern stingray (Dasyatis americana) swimming over sand in shallow water, split level photo with blue sky and clouds. The Sandbar, Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea
RF - Spinyhead blenny fish (Acanthemblemaria spinosa) extends out of its hole in a red sponge to snare passing food. Caribbean Sea
Longnose hawkfish (Oxycirrhites typus) hiding in coral, Tubbataha Reef Natural Park, UNESCO World Heritage Site, Sulu Sea, Cagayancillo, Palawan, Philippines
Verreauxs sifaka (Propithecus verreauxi) running across a road with bus in background, Berenty Private Reserve, southern Madagascar, August 2016
Asiatic elephants (Elephas maximus indicus) sniffing the air as they get close to a water body, Kaziranga National Park, Assam, India
Southern stingray (Hypanus americanus) at the surface. Stingray City, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea. Bait was used for this shot
Split level photo of an American crocodile (Crocodylus acutus) floating at the surface over a shallow seagrass meadow, close to mangroves
RF- Secretary blenny (Acanthemblemaria maria) peering from hole in massive Starlet coral (Siderastrea siderea). East End, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea
Adult female Fosa (Crytoprocta ferox) stalking prey on deciduous forest floor. Kirindy Forest, western Madagascar. November 2010
Coquerels sifaka (Propithecus coquereli) Palmarium, Atsinanana, Madagascar
Indri (Indri indri) feeding on fresh leaves / shoots in the rainforest canopy. Andasibe-Mantadia National Park, eastern Madagascar
Verreauxs sifaka (Propithecus verreauxi) female carrying baby, Berenty Private Reserve, Madagascar. Endangered species
Las Gralarias Glassfrog (Nymphargus lasgralarias) spawn with tadpoles developing in eggs hanging from leaf, Mindo, Pichincha, Ecuador
Pinocchio lizard (Anolis proboscis) male on stem, Mindo, Pichincha, Ecuador, Endangered species
Minute Glassfrog (Centrolene peristictum) male guarding eggs, Mindo, Pichincha, Ecuador, Vulnerable species
Mountain gorilla (Gorilla beringei beringei) female carrying baby on her back, member of Humba group, Virunga National Park, North Kivu, Democratic Republic of Congo, Africa, Critically endangered