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Fiordland crested penguins (Eudyptes pachyrhynchus) in dense coastal forest, Lake Moeraki, South Island, New Zealand, November, threatened species
Marine iguanas (Amblyrhynchus cristatus) silhouettes at sunset, Punta Vicente Roca, Isabela Island, Galapagos
The forests of the Modi Khola river valley shrouded in fog. Annapurna Sanctuary, central Nepal, November 2011
Japanese Macaques (Macaca fuscata) in hot springs, Jigokudani, Nagano Prefecture, Honshu, Japan
Darwins Orchid (Angraecum sesquipedale) species which is pollinated by a long-tongued moth, from Ambila, Madagascar. Photograph taken on location for BBC Wild Madagascar Series, August 2009
Nicaraguan giant glass frog (Espadarana prosoblepon) La Selva Field Station, Costa Rica
Green turtles (Chelonia mydas) swimming with angelfish, Punta Vicente Roca, Isabela Island, Galapagos
Great blue heron (Ardea herodias) taking off, with coastal landscape, Las Bachas, Santa Cruz Island, Galapagos
Resplendent quetzal (Pharomachrus mocinno) male, Talamanca mountains, Costa Rica
Starr Puffer (Arothron stellatus), juvenile swimming. Lembeh Strait, North Sulawesi, Indonesia
Rice (Oryza sativa) terraces. Jatiluwih Green Land, Bali, Indonesia. 2015
Palms growing in front of Rice (Oryza sativa) terrace. Jatiluwih Green Land, Bali, Indonesia. 2015
Coconut / Veined octopus (Amphioctopus marginatus) hunts in the sand at night, while illuminated by a divers torch beam, Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Pacific Ocean
Marine iguana (Amblyrhynchus cristatus) on shore, Punta Moreno, Isabela Island, Galapagos
Whooper swans (Cygnus cygnus) two in flight, during snowfall, Lake Kussharo, Japan, February
Backlit photo of a Robust ghost pipefish (Solenostomus cyanopterus) reveals a small fish that it has just eaten in its stomach. Lembeh Strait, North Sulawesi, Indonesia. Molluca Sea
Two Sulawesi black macaques / Celebes crested macaques (Macaca nigra) juveniles, in woodland, portrait, Tangkoko National Park, northern Sulawesi, Indonesia. Critically endangered
Three Red-crowned cranes (Grus japonensis) in flight. Hokkaido, Japan. March
Mimika bobtail squid (Euprymna morsei), Hokkaido, Japan, Pacific Ocean
Australian sea (Neophoca cinerea) juvenile playing in the sand, Carnac Island, Western Australia
Pelagic thresher shark (Alopias pelagicus) swimming in open ocean, Malapascua Island, Philippines, Pacific Ocean
Caspian Tern (Sterna caspia) in flight, diving for prey. Lake Ellesmere, Canterbury, South Island, New Zealand
Two Red-footed booby (Sula sula) perched on branches, Genovesa Island, Galapagos Islands, Ecuador
Lammergeier / Bearded vulture (Gypaetus barbatus) in flight with the out-of-focus moon behind. Ladakh, Himalayas, northern India
Australian sea lion (Neophoca cinerea) playful juvenile swimming upside-down, Carnac Island, Western Australia
A wide angle macro view of a Veined octopus ( Amphioctopus marginatus) stretching out from its protective den in an old clam shell. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Violet sabrewing hummingbird (Campylopterus hemileucurus) male hummingbirds territorial fight at dawn, Turrialba, Costa Rica
Black-crowned Central American squirrel monkey (Saimiri oerstedii) pair playing with each other, one with hand over eyes of other, Manuel Antonio National Park, Quepos, Costa Rica
Green turtle (Chelonia mydas) swimming near mangroves, Elizabeth Bay, Isabela Island, Galapagos
Madagascar green-emerald giant pill millipedes (Zoosphaerium neptunus) Andasibe-Mantadia National Park. Madagascar
Larval Wonderpus octopus (Wunderpus photogenicus) drifting in the open ocean at night off Anilao, Philippines
Polar bears (Ursus maritimus) standing up on hind legs, barrier island outside Kaktovik, Alaska, USA. September
Firefly squid (Watasenia scintillans) emitting light from photophores, Toyama Bay, Japan. This form of bioluminescence is used as counter illumination camouflage
Tasseled wobbegong shark (Eucrossorhinus dasypogon) yawning. Citrus Ridge, Yanggefo Island, Gam Island, Raja Ampat, West Papua, Indonesia
Ring-tailed lemur (Lemur catta) female carrying two babies. Anjaha Community Conservation Site, near Ambalavao, Madagasca
Reticulated glass frogs (Hyalinobatrachium valerioi) pair in amplexus, Osa Peninsula, Costa Rica
Male Blue-footed booby (Sula nebouxii) walking on sand, San Cristobal Island, Galapagos, South America
Snake-mimic caterpillar (Hemeroplanes triptolemus) a hawkmoth caterpillar that resembles a viper when threatened, on branch, La Selva Biological Station, Costa Rica
Male Mantled howler monkey (Alouatta palliata) on foot bridge, La Selva Biological Station, Costa Rica
Ocelot (Leopardus pardalis) climbing a tree trunk Costa Rica, Central America, 2016. Filmed for the BBC series Big Cats
Southern brown kiwi (Apteryx australis) primative flightless bird endemic to New Zealand, Stewart Island, New Zealand
Guineafowl puffer fish (Arothron meleagris) inflated, displaying defensive behaviour over coral reef, Yap, Indo Pacific, Micronesia
Great spotted kiwi (Apteryx hstii) Captive, Otorohanga Breeding Facility, New Zealand
Brown kiwi (Apteryx mantelli) in nocturnal kiwi house with reversed daylight cycle, Orana Wildlife Park, Christchurch, New Zealand. February
Short-snouted seahorse (Hippocampus hippocampus) male, Ponza Island, Italy, Tyrrhenian Sea, Mediterranean Sea
False tiger moth (Dysphania militaris) Isle of Marinduque, Philipines
Keel-billed toucan (Ramphastos sulfuratus) feeding, tossing fruit seed in beak, perched on branch. Boca Tapada, Costa Rica. Sequence 2/2
White-faced Capuchin (Cebus capucinus imitator) Carara National Park, Costa Rica
