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Sawfly larvae (Nematus pavidus) defensive posture, Peatlands Park, County Armagh, Northern Ireland. September
Land crab (Geosesarma sp. ) which raids Pitcher plant (Nepenthes ampullaria) for preyLand crab (Geosesarma sp.) which raids Pitcher plant (Nepenthes ampullaria) for prey, Sarawak, Borneo
Peacock mantis (Pseudempusa pinnapavonis) in defensive posture; captive occurs in Burma and Thailand
Shoebill stork (Balaeniceps rex) feeding on a Spotted African lungfish (Protopterus dolloi) in the swamps of Mabamba, Lake Victoria, Uganda
Calabar burrowing boa snake (Calabaria reinhardtii) in defensive ball, captive, occurs equatorial rain forest of West and central Africa. Head and tail are very similar
Trumpetfish (Aulostomus maculatus) using a Spanish Hogfish (Bodianus rufus) as a stalking horse. Spanish Hogfish is eating an arm of a starfish. Bonaire, Leeward Antilles, Caribbean
Pampas cat (Leopardus colocola) La Pampa Province, Argentina
Veined octopus (Amphioctopus marginatus) between clam shell halves. Ambon, Maluku Archipelago, Indonesia. Banda Sea, tropical west Pacific Ocean
Tawny nurse shark (Nebrius ferrugineus) group, individuals circling each other at dusk. Vavuu Atoll, Maldives. Indian Ocean
Northern goshawk (Accipiter gentilis) flying with squirrel prey, Finland. March
Blue tit (Cyanistes caeruleus) feeding young in the nestbox, Bavaria, Germany, May
Pheasant (Phasianus colchicus) male calling, East Frisian Islands, Wittbulten National Park. Wadden Sea UNESCO World Heritage Site, Germany, June
Lion (Panthera leo), females hunting buffalo, Masai-Mara Game Reserve, Kenya
Yellow-bellied toad (Bombina variegata) in defensive posture showing warning colours, on dried soil with mud cracks, Lower Saxony, Germany. August
Tiger (Panthera tigris), rolling on back and relaxing after feeding, Ranthambhore National Park, Rajasthan, India
Acrobat ant (Crematogaster sp) tending aphids on white oak (Quercus sp) Washington State Park, Pennsylvania, USA September
Pearlfish (Carapus acus) in anus of Sea cucumber (Stichopus regalis). These species have a commensal relationship, with the fish living in the gut of the sea cucumber. Catalonia, Spain
Frill-neck Lizard (Chlamydosaurus kingii), displays on a termite mound. Northern Territory, Australia
Humpback whale (Megaptera novaeangliae) breaching, Prince William Sound, Alaska, July
North American Bobcat (Lynx rufus) stalking along Madison River. Yellowstone National Park, Wyoming, USA. January
European flamingo (Phoenicopterus roseus) bathing Camargue, France, May
Black and white Jack-a-poo, Jack Russell cross Poodle, pup, 8 weeks old, jumping up
Grey tabby kitten standing and begging
Capercailie (Tetrao urogallus) male displaying from branch of Scots pine tree. Cairngorms National Park, Highlands, Scotland, UK, March
Barn swallow (Hirundo rustica) alighting at pond to collect mud for nest building, Scotland, UK, May
Prairie Rattlesnake (Crotalus viridis) sunbathing, Bozeman, Montana, USA
Juvenile of Amazon Tree-Boa (Corallus hortulanus) in defensive position. Yasuni National Park, Orellana, Ecuador
Mountain hare (Lepus timidus) grooming itself, with back foot raised, Cairngorms National Park, Scotland, UK, February. Highly Commended in the Animal Behaviour category of the British Wildlife
Striped mackerel (Rastrelliger kanagurta) mouth wide open as it swims through the water, filtering zoopankton with its gill rakers. Marsa Shouna, Port Ghalib, Marsa Alam, Egypt. Red Sea
RF - Sea otter (Enhydra lutris) floating, California, USA, July
Burrowing owl (Athene cunicularia) fledgling with head on side, Pantanal, Brazil. November
California sea lion (Zalophus californianus) uses a Panamic cushion star (Pentaceraster cumingi) as a toy. The sealions pick up the starfish and then drop them and chase after them as they sink
Green sea turtle female (Chelonia mydas) returning to the sea, Tortuguero National Park, Costa Rica
RF - Scarlet macaw (Ara macao) pair preening, Osa Peninsula, Costa Rica
Green-winged macaws (Ara chloroptera) preening each other. Brazil. South America
Reticulated giraffe (Giraffa camelopardalis reticulata) mother grooming baby, Samburu Game Reserve, Kenya, Africa, August
Atlantic salmon (Salmo salar) migrating for spawning in river, Gaspe Peninsula, Quebec, Canada, October
Arctic tern (Sterna paradisaea) in flight with fish in beak for feeding chick, Machias Seal Island, Bay of Fundy, New Brunswick, Canada, July
Brown throated three toed sloth (Bradypus variegatus) hanging from branch, Yasuni National Park, Orellana, Ecuador
Pod of Risso Dolphin (Grampus griseus) at surface, Baja California, Mexico
Sea otter (Enhydra lutris) female and pup playing. Alaska, USA, February
RF - Siberian tiger (Panthera tigris altaica) female grooming cub age three months, captive. (This image may be licensed either as rights managed or royalty free.)
Red Fox (Vulpes Vulpes) scratching in sunset light, North London, England, UK, June
RF - Cheetah (Acinonyx jubatus) playing with three cubs. Serengeti / Ngorongoro Conservation Area, Tanzania. (This image may be licensed either as rights managed or royalty free.)
Whale shark (Rhincodon typus) swimming through sun beams. This individual was following blue whales around to feed on their faeces. Indian Ocean, off Sri Lanka
Common swift (Apus apus) flying low to water while hunting, Norfolk, England, UK, July. Winner of Animal Behaviour Category of the British Wildlife Photography Awards 2019. Photographer
Tiger cardinalfish (Cheilodipterus macrodon) male mouth brooding a clutch of eggs behind his sharp teeth. Gubal Island, Egypt. Strait Of Gubal, Red Sea
Pin-tailed whydah (Vidua macroura) male in display flight. Allahein River, Gambia
