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Erect-crested penguin (Eudyptes sclateri) pair in greeting display. Antipodes Island, New Zealand Sub-Antarctic Islands. Endemic to Antipodes and Bounty Islands. Endangered species
RF- Gentoo Penguin (Pygoscelis papua) chick begging parent for food, Antarctica
Atlantic puffic, (Fratercula arctica) portrait. Machias Seal Island, Bay of Fundy, New Brunswick, Canada, August
American oystercatcher (Haematopus palliatus) feeding on Calico Clam, shortly after dawn, seashore, Tampa Bay, St. Petersburg, Florida, USA. April
Green-crowned brillian-t hummingbird (Heliodoxa jacula) visiting Torch ginger flower (Etlingera elatior), Poas Volcano National Park Costa Rica
Wrybill (Anarhynchus frontalis) standing shallow water with beak wide open. Lake Ellesmere, South Island, New Zealand. November
Golden eagle (Aquila chrysaetos) male head portrait, Kalvtrask, Vasterbotten, Sweden. December
Black-headed Ibis / Oriental Ibis (Threskiornis melanocephalus), Karnataka, India
Royal terns (Sterna maxima) pair crossing bills during courtship behaviour, Fort DeSoto Park, Florida, USA
Horned puffin (Fratercula corniculata) close-up portrait, St. Paul Island, Pribilofs, Alaska, USA, July
RF- Rufous Hummingbird (Selasphorus rufus) female feeding in flight on Scarlet Gilia (Ipomopsis aggregata). Gila National Forest, New Mexico, USA. August
RF- Capercaillie (Tetrao urogallus) male displaying, Inshriach, Cairngorms National Park, Scotland, UK. February
Greylag goose (Anser anser) taking flight from lake. Southern Norway. March
Red-crowned Crane (Grus japonensis) preening in snow, Hokkaido, Japan, February
Grasshopper Warbler (Locustella naevia) singing, Joensuu, Finland, May
Silvery-cheeked Hornbill (Bycanistes brevis) close-up at zoo, occurs in East Africa
Adult Helmet Vanga (Euryceros prevostii) incubating a clutch of 3 eggs. Nest constructed in the fork of a Birds Nest Fern (Asplenium sp.) Masoala National Park, north east Madagascar
Paradise Riflebird (Ptiloris paradiseus) young males size each other up between practice displays. Australia
Keel billed toucan (Ramphastos sulfuratus), captive
California condor (Gymnogyps californianus), IUCN Critically Endangered, captive
Greater flamingos (Phoenicopterus ruber roseus) males displaying, lake Nakuru, Kenya
Gannet (Morus bassanus) adult wing stretching and looking up, Shetland Islands, Scotland, UK, September
RF- Eagle owl (Bubo bubo) calling, raised in captivity and newly released into the wild. Stockholm archipelago, Uppland, Sweden. September
White tailed sea eagle (Halieetus albicilla) head portrait beak open calling, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary, January
Stellers Eagle (Haliaeteus pelagicus) close up of head and beak, Hokkaido, Japan, February
Ruppells weaver (Ploceus galbula) male building nest, Oman, May
Sword-billed Hummingbird (Ensifera ensifera) feeding at an Angels or Devil sSword-billed Hummingbird (Ensifera ensifera) feeding at an Angels or Devils Trumpet Flower (Datura sp.). Yanacocha montane cloud forest, Ecuador
Grey heron (Ardea cinerea) and Great egret (Ardea alba) fighting over fish, Lake Csaj, Pusztaszer, Hungary, December. Winner of the Portfolio category of the Terre Sauvage Nature Images Awards
Demoiselle crane (Anthropoides virgo) portrait, captive
Violet Sabrewing Hummingbird (Campylopterus hemileucurus) male hovering at feeder, Bosque de Paz, Costa Rica, Central America
Erect-crested penguins (Eudyptes sclateri) pair on coast, Proclamation Island, Bounty Islands, New Zealand Sub-Antarctic Islands. Endemic to Antipodes and Bounty Islands. Endangered species
RF- Gannets (Morus) displaying, Great Saltee, County Wexford, Ireland. May
Gannet (Morus bassanus) breeding pair during part of their elaborate courtship ritual. Saltee Islands, Republic of Ireland, May
Red-legged Seriema (Cariama cristata) portrait, captive, Apenheul Park, Netherlands
Dalmatian pelicans (Pelecanus crispus) low angle perspective of open bills, Lake Kerkini, Greece, February. Highly Commended, 2011 Wildlife Photographer of the Year competition
White throated kingfisher (Halcyon smyrnensis) perched on a reed, Sunderban tiger reserve, West Bengal, India
Kingfisher (Alcedo atthis) diving into water to catch fish, Near Bourne, Lincolnshire, England, UK. January
Blackbird (Turdus merula) male eating winfall apples in snow, Broxwater, Cornwall, UK. December
Two Great egrets (Ardea alba) face to face against white sky, Lake Csaj, Pusztaszer, Hungary. Winner of the Portfolio category of the Terre Sauvage Nature Images Awards competition 2015
Toco toucan (Ramphastos toco) head portrait, Pantanal wetlands, Mato Grosso do Sul, Brazil
Red and green macaw (Ara chloropterus) head portrait, Burraco das Araras, Pantanal wetlands, Mato Grosso do Sul, Brazil
Chestnut-eared aracari (Pteroglossus castanotis) portrait, Pantanal wetlands, Mato Grosso, Brazil
RF - Pied kingfisher (Ceryle rudis) female, perched on dead branch, Allahein River, The Gambia. (This image may be licensed either as rights managed or royalty free.)
Pair of Waved albatrosses (Phoebastria irrorata) billing as greeting. Punta Suarez, Espanola Island, Galapagos Islands, Ecuador
Keel-billed toucan (Ramphastos sulfuratus) perched in Trumpet tree (Cecropia peltata) feeding on fruit, Caribbean slope of Guatemala
Keel-billed toucan (Ramphastos sulfuratus) perched on branch with fruit in beak, Calakmul Bioshpere Reserve UNESCO Site, Campeche, Mexico
Ruddy duck (Oxyura jamaicensis) male in courtship display. Valkenhorst nature reserve, Valkenswaard, The Netherlands. May
Water rail (Rallus aquaticus) wading through water, Dumfries and Galloway, Scotland, UK
