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Praying mantis (Mantidae) with water vapour from cloud. Taken at high altitude hill station, Himalayas, India
Limestone microfossils in a 0. 03mm-thick slice of fossil-rich limestone viewed at high magnificationLimestone microfossils in a 0.03mm-thick slice of fossil-rich limestone viewed at high magnification, image area is 3mm across in real life
Western reef egret (Egretta gularis) wading in river at sunrise, Allahein River, The Gambia
Large school of juvenile Fusilier (Caesionidae) fish swimming in water column, Andaman Sea, Mu Koh Similan National Park, Phang-nga, Thailand
Beech (Fagus sp. ) trees in autumn, Saja Natural Park, Cantabria, Spain. NovemberBeech (Fagus sp.) trees in autumn, Saja Natural Park, Cantabria, Spain. November
Aerial view of the Izarilla River meandering across countryside lined with trees, Villaescusa, Cantabria, Spain. May
Reeftop pipefish (Corythoichthys haematopterus) swimming over hard coral (Diploastrea heliopora), Laamu Atoll, Maldives, Indian Ocean
Four Grey partridges (Perdix perdix) on snow-covered arable field, Scottish Borders, Scotland, UK. January
Wolf (Canis lupus) standing in snow, Yellowstone National Park, USA. January
Soft focus close up of male Banded demoiselle damselfly (Calopteryx splendens) body and wings, roosting among grasses. Cornwall, UK. July
Low angle view of Jaguar (Panthera onca) patrolling its territory at night, La Papalota, Nayarit, Mexico. Camera trap image
Silhouette of Axolotl (Ambystoma mexicanum) swimming in a woodland stream surrounded by towering trees, Sierra de Manantlan, Mexico
Svartifos waterfalls hiding wall of basalt columns, Vatnajokull National Park, Iceland. July
Two Mountain hares (Lepus timidus) running in deep snow, Monadhliath Mountains, Highlands, Scotland, UK. March
Turning European beech (Fagus sylvatica) leaves in autumn. Exmoor National Park, Somerset / Devon, England. November
Split level image of the tiny chapel of St Cwyfan, surrounded by sea at high tide and underwater forests of mixed seaweed near Aberffraw, Anglesey, Wales, Irish Sea, UK. July
Scalefin anthias / Scalefin basslet (Pseudanthias squamipinnis), in front of a gorgonian sea fan (Gorgonia sp)Scalefin anthias / Scalefin basslet (Pseudanthias squamipinnis), in front of a gorgonian sea fan (Gorgonia sp.), Fiji, Pacific Ocean
Eurasian spoonbill (Platalea leucorodia), portrait, Pusztaszer reserve, Hungary. May
Female Red fox (Vulpes vulpes) cautiously staring at the camera from the base of a moss-covered tree, Hungary. January
Honeycomb / Cushion starfish (Pentaceraster alveolatus) composite image on black background showing colour variations Malapascua Island, Philippines, Indo-Pacific species
Two male Alpine Ibex (Capra ibex) fighting on a mountain slope at dawn, Varaita Valley, Alps, Italy, July
Two Great white egret (Ardea alba) in winter, Kiskunsag National Park, Hungary. December
Grey heron (Ardea cinerea) and Great white egret (Ardea alba) standing facing each other at sunrise. Lake Csaj, Kiskunsagi National Park, Hungary. December
Three Lined seahorses (Hippocampus erectus), two male and one female, feeding on plankton in an alkaline saltwater pond, The Bahamas, Caribbean
Lily of the valley (Convallaria majalis) hidden at the bottom of the forest in spring. Morning sun is lighting from behind. Biebrza National Park, Poland
RF - Banded demoiselle damselfly (Calopteryx splendens) male roosting on reed, silhouetted against sun reflected in the water. Cornwall, England, UK. May
Sea star (Asterias rubens) on kelp, Vevang, Norway, Atlantic Ocean
Juvenile Mute swan (Cygnus olor) in early morning light, Valkenhorst Nature Reserve, The Netherlands, Europe. August
Common starling (Sturnus vulgaris) murmuration in between high voltage powerlines and electricity pylons, The Netherlands, Europe. February
Eastern horseshoe bat (Rhinolophus megaphyllus) flying out from an abandoned mine in late evening. Its orange colouring is due to bleaching from the intense ammonia atmosphere in the mine
Cranefly (Tipula paludosa) flight trail over meadow in old quarry on autumn night in Somerset, England. September
Saltpetre deposit in coastal lagoon, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, May, Digitally enhanced, removed dirt in lens. Bookplate
Small Stalked jellyfish (Calvadosia sp. ) attached to Bootlace weed / MermaidSmall Stalked jellyfish (Calvadosia sp.) attached to Bootlace weed / Mermaids hair (Chorda filum), Pendennis Head, Cornwall, UK. English Channel
Grey top-shell (Gibbula cineraria) grazing on Golden kelp (Laminaria ochroleuca). Pendennis Head, Cornwall, UK, English Channel
Water droplets on leaf creating a natural pattern, Tresco Tropical Garden, Tresco, Isle of Scilly, Cornwall, UK. June
Pair of Mute swans (Cygnus olor) at sunrise on misty lake. Poland, March
Snow covered trees in the Sotos del Alfaro Natural Reserve, Alfaro, La Rioja province, Spain. January
An Elegant starfish (Fromia elegans) on hard coral, Fiji
Red giant tajinaste / Mount Teide bugloss / Towers of jewels (Echium wildpretii) close up of flowers, Teide National Park, Tenerife, Canary Islands
Front view of an adult Black skimmer (Rynchops niger) in flight carrying a fish, Long Island, New York, USA. August
Wood pigeon (Columba palumbus) in falling snow. Leicestershire, UK, February
Northern gannets (Morus bassanus) pair displaying amongst campion, Bempton Cliffs, Yorkshire, UK, May
Orange tip butterfly (Anthocharis cardamines) male resting in dappled light created by dew droplets. The Netherlands. April
Sedge warbler (Acrocephalus schoenobaenus) singing with bokeh effect in background, Norfolk, England, UK, May. Winner of the Birds in Environment Category of the Bird Photographer of the Year 2015
Beech trees (Fagus sylvatica) and English bluebells (Hyacinthoides non-scripta) soft focus, West Wood, near Marlborough, Wiltshire, UK. May 2021
RF - Beech trees (Fagus sylvatica) and english bluebells (Hyacinthoides non-scripta). Late evening light and double exposure to create soft, dreamy effect. West Woods, nr Marlborough, Wiltshire, UK
Common spotted orchids (Dactylorhiza fuchsii), Kingcombe Meadows, Dorset, UK, June
Oxeye daisies (Leucanthemum vulgare)in upland hay meadow, Northumberland National Park, UK, June