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RF - Leopard (Panthera pardus) stalking prey, Londolozi Private Game Reserve, Sabi Sand Game Reserve, South Africa
Wild Mustang foal resting in wildflowers, Pryor Mountains, Montana, USA. June
Head portrait of wild dun roan Mustang mare in Sand Wash Basin, Colorado, USA
Wild rare Losino stallion standing in snow, Losa Valley, Burgos, Spain. January 2014
Bay Icelandic horse feeding in the snow, Snaefellsnes Peninsula, Iceland, March
Female Capybara (Hydrochoerus hydrochaeris) swimming in line with young after escaping a Jaguar attack (Panthera onca palustris) in a lagoon off the Paraguay River
Great crested grebe (Podiceps cristatus) close-up of an adult with two young chicks. The Netherlands. June 2014
Masai giraffe (Giraffa camelopardalis tippelskirchi) herd walking, Masai-Mara Game Reserve, Kenya. October
Adelie penguin (Pygoscelis adeliae) close up portrait, Antarctica
Sheep (Ovis aries) resting on the Quiraing mountains with low cloud cover, Isle of Skye, Inner Hebrides, Scotland, UK. July
Young rabbit hiding in grass, Okunoshima Rabbit island, Takehara, Hiroshima, Japan
White-tailed Eagle (Haliaeetus albicilla) in flight with mountains in background, Hokkaido, Japan, February
Wild Mustang horses fighting McCullough Peaks Herd Area, Wyoming, USA
Sperm whale (Physeter macrocephalus) tail fluke above water during dive, Kaikoura, New Zealand, July, Vulnerable species
Andean condor (Vultur gryphus) in flight over mountain peaks, Torres Del Paine National Park, Chile
RF- Eagle owl (Bubo bubo) calling, raised in captivity and newly released into the wild. Stockholm archipelago, Uppland, Sweden. September
Common / Eurasian crane (Grus grus) flock in flight with snow topped Pyrenees mountains in the background, during southward migration period, Haute Pyrenees, Gascony, France. December 2009
Young Polar bear (Ursus maritimus) walking across melting sea ice, Scott Inlet, Baffin Island, Canadian Arctic, August. Vulnerable species
Konik Wild Horse (Equus ferus caballus) yawning. The Netherlands, November
Common frog (Rana temporaria) sheltering from rain under toadstool (Macrolepiota procera) the Netherlands
Wild yak (Bos mutus) running, Kekexili, Qinghai, Tibetan plateau, China, December
Tibetan antelope (Pantholops hodgsonii) herd in the early morning with a low moon, Kekexili, Qinghai, Tibetan plateau, China, January
Wild horses / Mustangs, two stallions play fighting, Pryor Mountains, Montana, USA, July 2010
Wild female Amur leopard (Panthera pardus orientalis) on rocky hillside, Kedrovaya Pad reserve, Primorsky Krai, Far East Russia, January. Critically endangered species
Mustang / wild horses, second year cremello colt Claro play fighting with bay, McCullough Peak herd, Wyoming, USA, July 2009
King penguins (Aptenodytes patagonicus) on beach, South Georgia. Taken on location for BBC Frozen Planet series, 2008
Red Fox (Vulpes vulpes) vixen on a misty day in woodland, Black Forest, Germany, July. Runner-up in Animal Portraits category, Wildlife Photographer of the Year competition 2011
A wild newborn colt (Equus caballus) standing up being nuzzled by its mother. Foothills of Reno, Nevada, USA, May
Snowy Owl (Bubo scandiaca) in flight over snow covered landscape, Ottawa, Ontario, Canada, December
Adelie penguins (Pygoscelis adeliae) on iceberg, Antarctica, January
Group of lions (Panthera leo) at night, Masai Mara, Kenya. Photographed using Starlight Camera technology without artificial lighting
Two Cape Ground squirrels (Xerus inauris) greeting, Kgalagadi Transfrontier Park, South Africa Non-ex
Arctic tern (Sterna Paradisaea) hovering in flight over an iceberg. Peterman IslandArctic tern (Sterna Paradisaea) hovering in flight over an iceberg.Peterman Island, Antarctic Peninsula, Antarctica. February
Flock of Kelp / Southern black backed Gulls (Larus dominicanus) flying over iceberg. Graham Passage, Antarctic Peninsula, Antarctica. February. (digitally altered image)
Polar bear (Ursus maritimus) sow with her cub outside their den in late winter, Arctic coast of Alaska
Male Comet / Madagascar moon moth (Argema mittrei) recently emerged drying its wings. Andasibe-Mantadia National Park, Madagascar
Wild horse / mustang (Equus caballus) two young mustangs play-fighting in the Pryor Mountains, Wyoming, USA
Green woodpecker {Picus viridis} male in flight, Lorraine, France
Flock of Sanderling (Calidris alba) flying over a rough sea, Berwickshire Coast, Scotland, UK, March
Ice cliffs of the Ross ice shelf, Antarctica
Black browed albatross flying over iceberg {Thalassarche melanophrys} South Georgia
Tail fluke of a diving Humpback whale (Megaptera novaeangliae) misty coast in background, Alaska, USA. Gulf of Alaska, Pacific ocean
Fulmar (Fulmarus glacialis) rear view of bird hanging in air over steep cliffs, Shetland Islands, Scotland, UK. HIGHLY COMMENDED - Animals in their environment - Wildlife Photographer of the Year
RF- European hedgehog (Erinaceus europaeus) orphan, Jarfalla, Sweden. August
Amur / Siberian Tiger (Panthera tigris altaica) female in the wild, on a hillside, Lazovskiy zapovednik / Lazo Reserve protected area, Primorskiy krai, Far Eastern Russia, February 2012
Two wild Mustang stallions fighting in White Mountain Herd Area, Wyoming, USA. August
Wild Mustang stallion chasing another away from his family in White Mountain Herd Area, Wyoming, USA. August
RF- Marine Iguana (Amblyrhynchus cristatus) head portrait, Fernandina island, Galapagos, Ecuador