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Flock of Stellers eiders (Polysticta stelleri) in flight, Batsfjord, Norway, March
Commen raven (Corvus corax) flying in falling snow, Vardo, Norway, March
Arctic ground squirrel (Spermophilus parryii) portrait, Denali National Park, Alaska, USA, September
Male King eider (Somateria spectabilis) displaying, Norway, February
Arctic fox (Alopex lagopus) Svalbard, Norway, April
Spruce grouse (Falcipennis canadensis) in forest, Denali National Park, Alaska, USA, September
Pika (Ochotona princeps) resting, Denali National Park, Alaska, USA, September
Osprey (Pandion haliaetus) in flight with a fish, Finland, July
Brown bear (Ursus arctos) and cubs in the snow, Finland, April
Two juvenile Atlantic yellow legged gulls (Larus michahellis) in flight, Izmir, Turkey, October
Bald eagle (Haliaeetus leucocephalus) in flight, Alaska, USA, February
RF - Polar Bear (Ursus maritimus) males fighting, Churchill, Canada, November
RF - Reindeer (Rangifer tarandus) on a snowy ridge, Svalbard, Norway, April
Bison (Bison bison) on frost covered ground, Yellowstone National Park, USA, February
Grizzly Bears (Ursus arctos) resting, Lake Clarke National Park, Alaska, September
Grizzly Bear (Ursus arctos) asleep, Lake Clarke National Park, Alaska, September
Brown Bear (Ursus arctos) adult portait, Finland, June
Gannet (Morus bassanus) adult wing stretching and looking up, Shetland Islands, Scotland, UK, September
Bison (Bison bison) close up head portrait with snow on fur, Yellowstone National Park, USA, February
Black bears (Ursus americanus) three cubs, one climbing, Minnesota, USA, June
Coastal brown bear (Ursus arctos) female in a meadow, cubs nearby, Lake Clarke National Park, Alaska, USA, July
RF- Great grey owl (Strix nebulosa) in flight, Finland, March
RF- Razorbill (Alca torda) resting on rocks. Great Saltee Island, County Wexford, Ireland. April
Polar bear (Ursus maritimus) curious young male on the newly frozen pack ice, Beaufort Sea, off the 1002 area of the Arctic National Wildlife Refuge, North Slope, Alaska
Polar bear (Ursus maritimus) 3-year-olds play fighting on newly formed pack ice, Beaufort Sea, off the 1002 area of the Arctic National Wildlife Refuge, North Slope, Alaska
Pine marten (Martes martes) leaping, Ardnamurchan Peninsula, Scotland, August. Highly commended in the Animal Behaviour category of the BWPA (British Wildlife Photography Awards) Competition 2016
Great Spotted Woodpecker in snow (Dendrocopos major), Scotland, January
Atlantic Puffins (Fratercula arctica) three resting on a snow bank, Hornoya, Varanger, Norway, April
Black-throated diver (Gavia arctica), Finland, June
Tengmalm Owl (Aegolius funereus) perched on a branch, Finland
White tailed eagle (Haliaeetus albicilla) in flight with snowy background, Rausu, Japan, February
Stellers Eider (Polysticta stelleri) male swimming, Varangerfjord, Norway, April, Vulnerable species. Bookplate from Danny Greens The Long Journey North
Chamois (Rupicapra rupicapra), Ballons des Vosges Regional Natural Park, Vosges, Lorraine, France, December
Large white butterfly (Pieris brassicae), Vallee de l Eure (Eure Valley), Eure-et-Loir, France, August
Portrait of male Reindeer (Rangifer tarandus) with large antlers, in velvet. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, June
Lion (Panthera Leo) male, low angle shot, South Africa. Taken on location for Pride tv series
Common Swallowtail butterfly (Papilio machaon) backlit showing wing structure. Podere Montecucco, Orvieto, Umbria, Italy, October
Newly emerged adult Two-tailed Pasha butterfly (Charaxes jasius) Podere Montecucco. Orvieto, Umbria, Italy August
Lesser flamingos (Phoeniconaias minor) flock in Lake Bogoria, Rift valley, Kenya
Labiated jungle frog (Leptodactylus labrosus) portrait, Canande, Esmeraldas, Ecuador
Red kangaroo male bounding towards camera. {Macropus rufus) NSW, Australia. The kangaroo is the animal long jump champion, able to clear 13m (the length of a bus) from a standing start
Pyrenean chamois {Rupicapra rupicapra pyrenaica} Pyrenees, France
Strawberry poison frog (Oophaga pumilio) pair mating, Sarapiqui, Heredia, Costa Rica
Common / Green iguana (Iguana iguana) portrait, Sarapiqui, Heredia, Costa Rica
O Shaughnessys dwarf iguana / Red-eyed dwarf iguana (Enyalioides oshaughnessyi) portrait, Mashpi, Pichincha, Ecuador, Vulnerable species
Palmar tree frog (Boana / Hypsiboas pellucens) on plant stem, Mindo, Pichincha, Ecuador
Pebas stubfoot toad / Harlequin toad (Atelopus spumarius) stretching on branch, Yasuni National Park, Orellana, Ecuador, Vulnerable Species
Yellow-tongued / Dim anole (Anolis scypheus) on log, Yasuni National Park, Orellana, Ecuador
