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Common buzzard (Buteo buteo) flying, Cairngorms National Park, Scotland, UK, February
Mountain hare (Lepus timidus) feeding, close up, Scotland, UK, April
Merlin (Falco columbarius) female alighting onto perch, Glen Tanar, Cairngorms National Park, Scotland, UK, June
Black throated diver (Gavia arctica) swimming, Assynt, Sutherland, Highland, Scotland, UK, June
Buzzard (Buteo buteo) perched on alder snag, Cairngorms National Park, Scotland, UK, February
Golden eagle (Aquila chrysaetos) feeding on Red deer (Cervus elaphus) carcass, Assynt, Sutherland, Highland, Scotland, January
Sparrowhawk (Accipiter nisus) perching on lichen covered snag, Dumfries and Galloway, Scotland, UK, October
Red throated diver (Gavia stellata) on lochan in dawn mist, Cairngorms National Park, Scotland, UK, May
Brown hare (Lepus timidus) adult running across snow-covered field, Scotland, UK
Red squirrel (Sciurus vulgaris) on tree trunk, Cairngorms National Park, Scotland, UK, August
Jewel anemone (Corynactis viridis), North Uist, Outer Hebrides, Scotland, UK, May. Small repro only
European river otter (Lutra lutra) female bringing octopus ashore for cub, Shetland, Scotland, UK, October
A family of otters rest on the intertidal seaweed. European river otter (Lutra lutra) Shetland, Scotland, UK, July
Kestrel (Falco tinnunculus) female alighting onto perch, Scotland, UK
Greylag goose (Anser anser) taking flight from lake. Southern Norway. March
Great white shark (Carcharodon carcharias) Guadalupe Island or Isla Guadalupe, Pacific Ocean, Mexico, September. Vulnerable species
Zebu Cattle (Bos primigenius indicus), Los Llanos, Colombia, South America
Great crested grebe (Podiceps cristatus) close-up of an adult with two young chicks. The Netherlands. June 2014
Galapagos sea lion (Zalophus wollebaeki) emerging from water, Galapagos
Masai giraffe (Giraffa camelopardalis tippelskirchi) herd walking, Masai-Mara Game Reserve, Kenya. October
Marbled white butterflies (Melanargia galathea) visiting Greater knapweed (Centaurea scabiosa) flowers, Dorset, England. Digital Composite
Adelie penguin (Pygoscelis adeliae) close up portrait, Antarctica
Greenland shark (Somniosus microcephalus) with parasitic copepod (Ommatokoita elongata), under ice, Lancaster Sound, Nunavut, northern Baffin Island, Canada, Arctic
Black rhinoceros (Diceros bicornis) female and young in the mud, Masai-Mara Game Reserve, Kenya. February
Bull sharks (Carcharhinus leucas) in Beqa (Benga) Lagoon, a reserve where sharks are fed as part of a commercial diving operation. Viti Levu, Fiji, South Pacific
Close up of European dark bee (Apis mellifera mellifera - nigra race), feeding on ice plant flowers (Sedum spectabile), Monmouthshire, Wales, UK. September
Red-crowned Crane (Grus japonensis) preening in snow, Hokkaido, Japan, February
Grasshopper Warbler (Locustella naevia) singing, Joensuu, Finland, May
Capercaillie (Tetrao urogallus) male in flight, Jalasjarvi, Finland, April
Black Grouse (Tetrao tetrix) male displaying with breath condensing in cold air, Kuusamo, Finland April
Young rabbit hiding in grass, Okunoshima Rabbit island, Takehara, Hiroshima, Japan
Rear view of Wildebeest (Connochaetes taurinus) herd crossing the Mara river, Masai-Mara game reserve, Kenya
Spotted reed frog (Hyperolius pucticulatus) tadpoles, captive, endemic to Tanzania
European hornet (Vespa crabro) worker in flight, Surrey, England, August
White-tailed Eagle (Haliaeetus albicilla) in flight with mountains in background, Hokkaido, Japan, February
Coalition of male Lions (Panthera leo) in grassland before storm, Masai-Mara game reserve, Kenya
Blue shark (Prionace glauca) close up, Azores, Portugal. Finalist in Wildlife Photographer of the Year Awards (WPOY) Competition 2014
Field of Giant plumrose anemones (Metridium giganteum) with a Snakelock anemone / Crimson sea anemone (Cribrinopsis fernaldi) in foreground, Alaska, USA, Gulf of Alaska. Pacific ocean
Sea otter (Enhydra lutris) floating on its back at surface among kelp, Alaska, USA. Gulf of Alaska, Pacific ocean
Detail of the pectoral fin of Roundhead parrotfish (Scarus viridifucatus) coast of Dhofar and Hallaniyat islands, Oman. Arabian Sea
Marine iguana (Amblyrhynchus cristatus) swimming back to the surface, Galapagos. Pacific ocean
Whale shark (Rhincodon typus) with shoal of anchovies staying near to avoid predation. Los Roques, Venezuela. Caribbean
Ground squirrels (Xerus inauris ) one with its tail over the back of another, Kgalagadi Transfrontier Park, Northern Cape, South Africa, February
Harlequin sweetlips (Plectorhinchus chaetodonoides) with cleaning Bluestreak cleaner wrasse (Labroides dimidiatus) and Bicolour cleaner wrasse (Labroides bicolor) South Atoll, Tubbataha Atolls
Musk ox (Ovibos moschatus) two covered in snow, Wrangel Island, Far Eastern Russia, September
Pair of Blesbok (Damaliscus pygargus) galloping across the open flood plains of the Bushmans River, Kariega Game Reserve, South Africa. Overall winner of the GDT Competition 2014
Fallow deer (Dama dama) buck crossing road under street lights. London, UK. January
Greylag geese (Anser anser). Cley, Norfolk, UK, March. Digital composite