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Gloucester old spot piglets (Sus scrofa domestica) suckling at the Cotswold Farm Show, Cirencester, UK, July 2008
Rothschilds giraffe (Giraffa camelopardalis rothschildi) close up of young calf skin pattern, captive
Grevy's zebra (Equus grevyi) close up, captive
Chinese Crested Dog, hairless bitch with puppy, male, 4 months
Common buckeye butterfly (Junonia coenia) feeding on Ironweed (Vernonia altissima) flower, North Guilford, Connecticut, USA, August
Spotted Hyaena (Crocuta crocuta) rolling in Devils Thorn flowers (Tribulus terrestris). Etosha National Park, Namibia, January
Great Tit (Parus major) in flight. Bavaria, Germany, December
Rough coated Jack Russell Terrier puppy, black, tan and white, portrait
Domestic dog, Old English Sheepdog / Bobtail, studio portrait
Pronghorn Antelope (Antilocapra americana) head portrait, South Dakota, USA
Large adult male Leopard (Panthera pardus) walking through the bush at night, Sabi Sand Private Game Reserve, South Africa, June
Close up of coiled tail of Parsons chameleon {Calumma parsonii} showing scales and strong green colouration. Tropical rainforest, Masoala Peninsula National Park, north east Madagascar
Snowy Owl (Bubo scandiaca) in flight over snow covered landscape, Ottawa, Ontario, Canada, December
Two Cheetahs (Acinonyx jubatus) touching noses in greeting display, Serengeti NP, Tanzania, East Africa, January
Bobtail / Old English Sheepdog, sitting alongside a longhaired Chihuahua, tan and white coated
Violet Sabrewing Hummingbird (Campylopterus hemileucurus) male hovering at feeder, Bosque de Paz, Costa Rica, Central America
Roan antelope (Hippotragus equinus) with young offspring, Mlilwane nature reserve breeding programme, Swaziland, Southern Africa
Two Cape Ground squirrels (Xerus inauris) greeting, Kgalagadi Transfrontier Park, South Africa Non-ex
Head portrait of Sumatran tiger (Panthera tigris sumatrae) with face half cast in shadow, with aggressive expression, captive
King Penguin (Aptenodytes patagonicus) adult standing amongst colony of chicks, Gold Harbor, South Georgia, South Atlantic Islands
Gannet (Morus bassanus) breeding pair during part of their elaborate courtship ritual. Saltee Islands, Republic of Ireland, May
Raccoon (Procyon lotor) standing up investigating camera, portrait, Stanley park, Vancouver, British Columbia, Cananda, September
Baby ground squirrel (Xerus inauris) using its tail for shade, Kgalagadi Transfrontier Park, South Africa. Non-ex
RF- Juvenile Sumatran tiger (Panthera tigris sumatrae), aged four months, captive, occurs in Sumatra, Indonesia
Tiger (Panthera tigris tigris), male portrait, Bandhavgarh, India, February 2013
Bengal Tigers (Panthera tigris tigris) adult female drinking water, her juvenile cub beside her. Bandhavgarh National Park, India, April
Bengal Tiger (Panthera tigris tigris) captive, with leaves digitally added
Bengal Tiger (Panthera tigris) sub-adult, approximately 17-19 months old, amongst forest vegetation. Endangered. Bandhavgarh National Park, India. Non-ex
Portrait of Siberian Husky dog face used as sled dogs inside Riisitunturi National Park, Lapland, Finland
Amur / Siberian Tiger (Panthera tigris altaica) female in the wild, on a hillside, Lazovskiy zapovednik / Lazo Reserve protected area, Primorskiy krai, Far Eastern Russia, February 2012
RF- Adult male Serval (Felis serval) stalking prey in long grass. Serengeti National Park, Tanzania
Young, female Grey seal (Halichoerus grypus) swimming over a forest of Cuvie kelp (Laminaria hyperborea), Inner Hebrides, Scotland, UK, Atlantic Ocean. August
Male Jaguar (Panthera onca) walking through rainforest, Belize, Central America, 2017. Captive. Filmed for the BBC series Big Cats
Black and white ruffed lemur (Varecia variegata) hanging from tree, Madagascar, critically endangered
RF- Morpho butterfly displaying on leaf (Morpho achilles). Amazonia, Ecuador
RF- Red Admiral butterfly (Vanessa atalanta) perched on Bigtooth Maple (Acer grandidentatum). Lost Maples State Park, Hill Country, Central Texas, USA. November
Marsh fritillary butterfly (Euphydryas aurinia) feeding on Wild garlic / Ramsons (Allium ursinum) at Strawberry Banks, Gloucestershire Wildlife Trust (GWT), Nature Reserve, Gloucestershire, UK. May
Close-up of scales of the Yellowbar Angelfish (Pomacanthus maculosus). Red Sea, Egypt
RF - Spicebush swallowtail butterfly (Papilio troilus) nectaring on Zinnia in farm garden, wild and free. Madison, Connecticut, USA
RF - Monarch butterfly (Danaus plexippus) with life-cycle tied to the presence of milkweed plants (Asclepias sp). East Haddam, Connecticut, USA. September
Marbled white butterfly (Melanargia galathea) resting on grass head, backlit against black background, Dunsdon, Devon, UK. July 2014
Duke of Burgundy butterfly (Hamearis lucina), Wiltshire, UK, June
Scarce tortoiseshell butterfly (Nymphalis xanthomelas) on pussy willow, Luhanka, Keski-Finland, April. / Central Finland, Finland, April
Comma butterfly (Polygonia c-album) feeding on wild aster, Sutcliffe Park Nature Reserve, Eltham, London, UK, September
Emperor Moth (Saturnia pavoniella) male, on tree bark, Orvieto, Umbria, Italy, April
Close-up of Swallowtail (Papilo machaon) wings, Emilia Romagna Region, Italy
Oriental sweetlips (Plectorhinchus vittatus), Maldives, Indian Ocean
RF- Pinto Quarter horse in snow at ranch, Shell, Wyoming, USA, February