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Bullethead parrotfish (Chlorurus sordidus) male, close up of eye, whilst sleeping at night. The Alternatives, Sinai, Egypt. Red Sea
Pallass cat (Otocolobus manul) Tibetan Plateau, Qinghai, China
Pallass cat (Otocolobus manul) peering, Tibetan Plateau, Qinghai, China
Reticulate stargazer (Uranoscopus sp. ) portrait. Ambon Bay, Ambon, Maluku Archipelago, IndonesiaReticulate stargazer (Uranoscopus sp.) portrait. Ambon Bay, Ambon, Maluku Archipelago, Indonesia. Banda Sea, tropical west Pacific Ocean
Golden eagle (Aquila chrysaetos) male head portrait, Kalvtrask, Vasterbotten, Sweden. December
Desert monitor (Varanus griseus griseus) occurs in Sahara Desert. Controlled conditions
Tricolour merle Collie puppy, Indie, age 10 weeks
Blue shark (Prionace glauca) close up, Azores, Portugal. Finalist in Wildlife Photographer of the Year Awards (WPOY) Competition 2014
Harlequin sweetlips (Plectorhinchus chaetodonoides) with cleaning Bluestreak cleaner wrasse (Labroides dimidiatus) and Bicolour cleaner wrasse (Labroides bicolor) South Atoll, Tubbataha Atolls
Yacare Caiman (Caiman yacare) with butterfly (Paulogramma pyracmon) resting on its snout, Pantanal, Brazil
African horned viper (Cerastes cerastes), captive
Unau / two-toed sloth (Choloepus didactylus) portrait, French Guiana
Lemon sole (Microstomus kitt) portrait. Gardur, south west Iceland. North Atlantic Ocean
Bison (Bison bison) close up head portrait with snow on fur, Yellowstone National Park, USA, February
Smooth-fronted caiman / Crowned dwarf caiman (Paleosuchus trigonatus) head above water, Cuyabeno, Sucumbios, Ecuador
Black caiman (Melanosuchus niger) at water surface with horse flies on its head, Yasuni National Park, Orellana, Ecuador
Parsons Chameleon (Calumma parsonii) close up of eye and skin, Andasibe-Mantadia National Park, Moramanga, Madagascar
Clown frog (Dendropsophus leucophyllatus), Villa Carmen Biological Station, Peru
Horned adder (Bitis caudalis) close up of eye, Brandberg area, Namibia
Clarion Burrowing Owl (Athene cunicularia rostrata), Clarion Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site (Socorro Islands)
Jumping spider (Salticidae) hunting among vegetation, San Jose, Costa Rica
Painted lady butterfly (Vanessa cardui) on flower, Peak District National Park, Derbyshire, UK. September
Emerald damselflies (Lestes sponsa) mating, with male above. Peak District National Park, Derbyshire, UK. September
RF- Common frog (Rana temporaria) portrait, Cornwall, UK, January
Eyes of Margined sole fish (Brachirus heterolapis) on its camouflaged body. Saonek Island, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean. Dampier Strait
Reticulate stargazer (Uranoscopus sp. ) emerging from the sandReticulate stargazer (Uranoscopus sp.) emerging from the sand. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Butterfish (Pholis gunnellus) portrait. Gardur, south west Iceland. North Atlantic Ocean
Red Irish lord (Hemilepidotus hemilepidotus) hides on the sebed, with Quillback rockfish (Sebastes maliger) and copper rockfish (Sebastes caurinus) behind in the bull kelp forest
Secretary blenny (Acanthemblemari maria) yawns as it peers out from a hole in the reef. West Bay, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
Coleman shrimp (Periclimenes colemani) close up, on a fire urchin (Asthenosoma varium). Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
Male Bullethead parrotfish (Chlorurus sordidus) displaying in late afternoon. Ras Katy, Sinai, Egypt. Gulf of Aqaba, Red Sea
RF - Siberian tiger (Panthera tigris altaica) in snow, captive
Red fox (Vulpes vulpes) close-up of half of face, captive
Demoiselle crane (Anthropoides virgo) portrait, captive
Portrait of Waglers / Temple Pitviper (Tropidolaemus wagleri) showing the thermo-receptive pit organ between the eye and mouth. Controlled conditions. Mindanao, Philippines, February
Black Jumping spider {Salticidae}. Tropical rainforest, Masoala Peninsula National Park, north east Madagascar
Fangtooth (Anoplogaster cornuta) portrait, Mid-Atlantic Ridge, North Atlantic Ocean
Pacific hatchetfish (Argyropelecus affinis) portrait
Two newly fledged burrowing owl chicks (Athene cunicularia) one being groomed by its mother (far left) Pantanal, Brazil. WINNER
Horned adder (Bitis caudalis) eyes and horns, Swakopmund, Erongo, Namibia
Close-up of an eye of a Sumatran tiger (Panthera tigris sumatrae), captive, occurs in Sumatra, Indonesia
Bengal Tiger (Panthera tigris tigris) captive, with leaves digitally added
Three Common toads (Bufo bufo) in amplexus, Lucerne, Switzerland. March. Cropped
Jumping spider (Heliophanus sp. ) portrait, Lucerne, Switzerland. AprilJumping spider (Heliophanus sp.) portrait, Lucerne, Switzerland. April
Jumping spider (Hypositticus pubescens) male, resting on leaf, Lucerne, Switzerland. June. Focus stacked
Small pearl-bordered fritillary butterfly {Boloria selene} head-on view, resting on grass, Cornwall, UK. June
Rusty whipsnake / Wagler's sipo (Chironius scurrulus) coiled up, close up, Yasuni National Park, Orellana, Ecuador
Swallow-tailed gull (Creagrus furcatus) eye detail, San Cristobal Island, Galapagos National Park, Galapagos Islands