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Moose (Alces alces) bull swimming across river. Grand Teton National Park, Wyoming, USA. September
RF - American bison (Bison bison) male standing near Punch Bowl thermal pool, green algae in foreground. Yellowstone National Park, USA
North American Bobcat (Lynx rufus) stalking along the edge of the Madison River. Yellowstone National Park, Wyoming, USA. January
Humpback whales (Megaptera novangliaea) pod engaged in social foraging by herding herring and other fish with bubble net feeding technique, Chatham Strait, Alaska, USA July
Eared grebes (Podiceps nigricollis) stand erect in water during courtship, North Park, Colorado, USA. June
Desert bighorn sheep (Ovis canadensis nelsoni) rams on steep sandstone wall. Valley of Fire State Park, Nevada, USA. February
White-tailed deer (Odocoileus virginianus) doe and fawn standing amongst Bulrushes on snow covered pond. Acadia National Park, Maine, USA. January
Black bear (Ursus americanus), two cubs playing, climbing up tree trunk. Yellowstone National Park, Wyoming, USA. June
American bison (Bison bison) female grazing near thermal pool, snow on ground. Biscuit Geyser Basin, Yellowstone National Park, USA. February 2019
American bison (Bison bison), two females covered in hoar frost near hot spring, portrait. Midway Geyser Basin, Yellowstone National Park, USA. February
Morning Glory thermal pool surrounded by snow, coniferous forest in background. Near Daisy Geyser Basin, Yellowstone National Park, USA. January 2019
Grizzly bear (Ursus arctos) running through water, Lake Clark National Park, Alaska, September
Moss-covered Sitka spruce (Picea sitchensis) and sword fern (Polystichum munitum) in the Hoh Rain Forest, Olympic National Park, Washington, USA, June
Frost on soaptree yuccas (Yucca elata) and winter morning fog, in the gypsum sand dunes of White Sands National Monument, New Mexico, USA. January
Flannel bush (Fremontodendron California Glory ) exposed and overflowing nectaries fluorescuing in UV light. Native to southwest USA
Southern magnolia / Bull bay (Magnolia grandiflora) flower. The opening and closing of petals controls the time beetle pollinators enter and leave the flower
Texas bluebonnets (Lupinus texensis) and Indian paintbrush (Castilleja coccinea) amongst wildflowers in ditch beside byway. Rains triggered germinaton of annual seeds. Cuero, Texas, USA. April
Tulip tree (Liriodendron tulipifera) flower, pollen and nectar fluorescing in UV light. Surrey, England, UK. Native to USA. Focus stacked. Sequence 2/2
Southern catalpa (Catalpa bignonioides) flowers. Recently opened with yellow nectar guides, purple in older pollinated flowers. Surrey, England, UK. Native to USA
Buacaille Etive Mor and River Etive in winter, Rannoch Moor, Glencoe, Scotland, UK.December
North American porcupine (Erethizon dorsatum) in snow, Vermont, USA. (Habituated rescued individual returned to the wild)
Great horned owl (Bubo virginianus) preening two chicks in nest in Saguaro (Carnegiea gigantea) cactus, Sonoran desert, Arizona, USA
American alligator (Alligator mississippiensis) emerging from water, in evening light. Myakka River State Park, Florida, USA. March
Whale shark (Rhincodon typus) close to the camera with a golden trevally fish (Gnathanodon speciosus) and outline of another whale shark behind, in the plankton-rich green waters of La Paz bay
Berkshire piglet in a basket among squashes in early autumn; Rhode Island, USA. October
Beaver pond with beaver lodge (on the left side) and trees reflected in autumn. Acadia National Park, Maine, USA. October
Moose (Alces alces) bull, crossing mountain river after sunset. Grand Teton National Park, Wyoming, USA. October 2017
Shoreline of Lake Powell, seen from Alstrom Point on Romana Mesa. Glen Canyon National Recreation Area, on the Arizona/Utah border, USA, January
Snowy egret (Egretta thula) shows its breeding plumage. St. Johns Management Area, Florida, USA, March
Western screech owl (Megascops kennicottii) is camouflaged as it looks from a tree cavity. Bosque del Apache National Wildlife Refuge, New Mexico, USA, January
Soaptree yucca (Yucca elata) at sunset. White Sands National Monument, New Mexico, USA. January
Sea Otter, (Enhydra lutris), young male, Elkhorn Slough, California, USA. January
Arctic tern (Sterna paradisaea), two fighting in mid-air. Machias Seal Island, New Brunswick / Maine, Canada / USA. July
American oystercatcher (Haematopus palliatus) adult feeding chick, Long Island, New York, USA. June
American avocets (Recurvirostra americana) two during aggressive (territorial) encounter, Bear River Migratory Bird Refuge, Utah, USA, May
Common loon (Gavia immer) adult in breeding plumage, preening in the last light of day, Michigan, USA, June
Blue jay (Cyanocitta cristata) holding an acorn in its bill whilst perched on tree stump, with autumn foliage, New York, USA, October
Black skimmers (Rynchops niger), three in non-breeding plumage on beach, Fort De Soto Park, St. Petersburg, Florida, USA. January. Digitally altered to remove distractions
White pelicans (Pelecanus erythrorhynchos) at Bear River Migratory Bird Refuge with an approaching storm in May, Utah, USA, May
Yellow-headed Blackbird (Xanthocephalus xanthocephalus) male perched in marsh vegetation (Phragmites) in foreground. Bear River Migratory Bird Refuge, view toward east
Indigo Bunting (Passerina cyanea) male in breeding plumage perched in Crabapple (Malus sp.) blossom in spring, Ithaca, New York, USA, May
Loggerhead shrike (Lanius ludovicianus) on an ice-covered fence after an ice storm, Anahuac National Wildlife Refuge, Texas, USA. January
Pileated Woodpecker (Dryocopus pileatus) male, at feeding excavation in tree trunk in winter, Freeville, New York, USA, January
Pileated woodpecker (Dryocopus pileatus) male excavating in search of food, New York, USA. Digital composite
Red-breasted Mergansers (Mergus serrator) male in foreground performing courtship display to female in background (left), Aurora, New York, USA, April
Abstract patterns of reflected sky and clouds on the ocean with a silhouetted Red-faced cormorant (Phalarocorax urile) in flight, St. Paul Island, Pribilof Islands, Alaska, USA, July
Common loons (Gavia immer), two adults and chick swimming on a misty lake in early morning, Michigan, USA. June
