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Grizzly cubs (Ursus arctos) two play fighting in the mist, near Brooks Falls in Katmai National Park, Alaska, USA, July
Grizzly Bear (Ursus arctos) resting, Lake Clarke National Park, Alaska, September
California sea lions (Zalophus californianus) playing in a kelp forest off Santa Barbara Island, California, USA
RF - Coyote (Canis latrans) in snow, Yellowstone. February
Red fox (Vulpes vulpes) resting and sitting on snow, Yellowstone National Park, USA, February
Coastal brown bear (Ursus arctos), Lake Clarke National Park, Alaska, September
Brown Bear (Ursus arctos) mother and 3-4 month cubs foraging for clams. Katmai National Park, Alaska, USA, June
Black bear (Ursus americanus) mother with cub. Yellowstone National Park, Wyoming, USA, May
Coastal brown bear (Ursus arctos) fishing, Lake Clarke National Park, Alaska, September
Porcupine (Erethizon dorsatum) on tundra. North Slope, Alaska, USA. September
Coyote (Canis latrans) in wintry landscape, Yellowstone National Park, USA, February
Coastal brown bear (Ursus arctos) and cub, Lake Clarke National Park, Alaska, September
Bighorn Sheep (Ovis canadensis) on the hillside, ruminating, Yellowstone National Park, Wyoming, USA. January 2011
Coyote (Canis latrans) concealed in the snow, Yellowstone National Park, Wyoming, USA. January
Red fox (Vulpes vulpes) in snow, Yellowstone National Park, Wyoming, USA. Wyoming, USA, January
Bison (Bison bison) standing steam from geothermal springs, Yellowstone National Park, Wyoming, USA, February
RF- Bison (Bison bison) standing in steam from geothermal springs. Yellowstone National Park, Wyoming, USA, February
RF- Very young Florida manatee (Trichechus manatus latirostrus) Manatees are often born with very wrinkled skin. Crystal River, Florida, USA. February 2010
Yellow warbler (Dendroica petechia) male singing. Bozeman, Montana, USA
White-winged Dove (Zenaida asiatica), adult perched on icy branch of Yaupon Holly (Ilex vomitoria), Hill Country, Texas, USA. February
Bald eagle (Haliaeetus leucocephalus) in flight, Homer, Alaska, USA, March
RF - Great horned owl (Bubo virginianus) in woodland, captive; Alaska, USA
Western Scrub-Jay (Aphelocoma californica), adult calling on icy branch of Possum Haw Holly (Ilex decidua) with berries, Hill Country, Texas, USA. February
Short-billed Dowitcher (Limnodromus griseus) on mudflats. Grays Harbor County, Washington. May
Barred Owl (Strix varia) fledgling resting in tree, King County, Washington, USA May
Spotted Owl (Strix occidentalis). Willamette National Forest, Oregon. June
Sandhill Cranes (Grus canadensis) feeding in agricultural fiields during migration. Central Nebraska, USA. March
Spruce grouse (Dendragapus fuliginosus) stops while foraging among fall blueberry bushes. Denali National Park, Alaska, USA
American oystercatcher (Haematopus palliatus) feeding, Fort Myers Beach, Gulf Coast, Florida, USA, March
Horned puffins (Fratercula corniculata) pair interacting by touching bills while perched on cliff ledge, St. Paul Island, Pribilofs, Alaska, USA, July
Great egret (Ardea alba) in flight, Florida, USA, March
Lazuli Bunting (Passerina amoena) perched, Grand Teton National Park, Wyoming, USA, June
Burrowing owl (Athene cunicularia) Adams County, Washington, USA. May
White Ibis (Eudocimus albus) feeding off Fort Myers beach, Gulf Coast, Florida, USA, March
Common merganser (Mergus merganser) male taking off from water, New York, USA, April
Brown pelicans (Pelecanus occidentalis) on water, split level view, Galapagos
Lower Falls at Artist Point, Yellowstone River. Yellowstone National Park, Wyoming, USA, June
Polar bear (Ursus maritimus) sow with two juveniles walk along the ice edge during autumn freeze up, Beaufort Sea, off Arctic coast, Alaska
White ibis (Eudocimus albus) flock in breeding plumage, backlit against dark background, four individuals flying in to join flock, Tampa Bay, Florida, USA, March
Grey wolf (Canis lupus) Katmai National Park, Alaska, USA, August
Rock formations silhouetted at sunset on the Pacfic coast of Olympic National Park, Washington State, USA
Wild Mustang pinto foal nuzzling up to mother, Sand Wash Basin Herd Area, Colorado, USA
Wild Mustang foals among wild flowers, Pryor Mountains, Montana, USA
White-tailed deer (Odocoileus virginianus) in snow, Acadia National Park, Maine, USA, February
Mallard (Anas platyrhynchos) walking on ice, Acadia National Park, Maine, USA, April
Icicles hanging from sandstone cliffs on shoreline, Apostle Islands National Lakeshore, Lake Superior, Squaw Bay, Wisconsin, February 2014
Water rushing down alpine stream, Logan Pass, Glacier National Park, Rocky Mountains, Montana, July 2010
Grey wolf (Canis lupus) portrait, Katmai National Park, Alaska, USA, August