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Hanging thief robber fly (Diogmites sp) Philadelphia, Pennsylvania, USA, July
Steller sea lions (Eumetopias jubatus) playing underwater, Inian Island, Alaska, USA. August
American red squirrel (Tamiasciurus hudsonicus) on tree trunk drinking in a beaver pond early in the morning. Acadia National Park, Maine, USA. May
Ridgeways rail (Rallus obsoletus levipes) foraging in wetlands, Bolsa Chica Ecological Reserve, California, USA April
California least terns (Sternula antillarum browni) courtship behaviour, Bolsa Chica Ecological Reserve, California, USA June
Blue jays (Cyanocitta cristata) confrontation in snow, April 2020. New York State, USA
Northern Cardinals (Cardinalis cardinalis) male and female in the snow, photographed from low angle, Freeville, New York, USA. April 2020
Yellow Warbler (Setophaga petechia, formerly Dendroica petechia), male in breeding plumage, Ithaca, New York, USA. May 2020
Canada goose (Branta canadensis) with gosling, a few days old. Massachusetts, USA. April
Wood duck (Aix sponsa) male in breeding plumage. Acadia National Park, Maine, USA. April
Long-beaked common dolphin (Delphinus capensis) off the coast of California, United States
Common Merganser (Mergus merganser) female with ducklings, in threat posture at the approach of a second female, Lansing, New York, USA
Baltimore Oriole (Icterus galbula) male with spider prey, perched in crabapple blossom in spring, Ithaca, New York, USA
Purple Finch (Carpodacus purpureus) perched amid apple blossom, New York, USA
Greater Roadrunner (Geococcyx californianus), sunbathing. Warming itself by erecting feathers to allow sun to strike directly on black skin. Arizona, USA
Ring-billed Gull (Larus delawarensis) perched on a No Fishing sign, New York, USA
Spotted Salamander (Ambystoma maculatum) in early spring migration to woodland pond, New York, USA
Common loon (Gavia immer), chick on lake, Michigan, USA. June
Sheephead (Semicossyphus pulcher) female, and Garibaldi, (Hypsypops rubicundus) in forest of giant kelp (Macrocystis pyrifera) off Santa Barbara Island, California, USA
Soaptree yucca (Yucca elata) on gypsum dun, White Sands National Monument, New Mexico, USA. December
RF - Petrified trees and travetine cascade. Geothermal feature at Mammoth Hot Springs. Yellowstone National Park, USA. (This image may be licensed either as rights managed or royalty free.)
RF - Red fox (Vulpes vulpes) in deep snow. Yellowstone National Park, USA. January (This image may be licensed either as rights managed or royalty free.)
RF - North American river otters (Lutra canadiensis) on the frozen river edge. Yellowstone National Park, USA. January (This image may be licensed either as rights managed or royalty free.)
Coyote (Canis latrans) foraging in deep winter snow disturbed by grazing American bison (Bison bison) Hayden Valley, Yellowstone National Park, Wyoming, USA. January
North American river otters (Lutra canadiensis) probaly female with two juveniles, on the frozen river edge. Upper Yellowstone River, Hayden Valley, Yellowstone, USA. January
RF - Two Grizzly bears (Ursus arctos) Lake Clark National Park, Alaska, September. (This image may be licensed either as rights managed or royalty free.)
North American river otter (Lutra canadiensis) two on frozen river edge. Upper Yellowstone River, Hayden Valley, Yellowstone National Park, Wyoming, USA. January
Coyote (Canis latrans) foraging in snow. Hayden Valley, Yellowstone National Park, USA. January 2019
American Bison (Bison bison) frost covered male, standing by dead trees. Firehole River Valley. Yellowstone National Park, Wyoming, USA. January
Rocky mountain bighorn sheep (Ovis canadensis canadensis) grazing. Lamar Valley, Yellowstone National Park, Wyoming, USA. January
RF - Herd of American Bison (Bison bison) around geo-thermal features. Firehole River Valley. Yellowstone National Park, Wyoming, USA. January 2008
Sun beam at sun rise over the Grand Canyon of the Yellowstone, Yelllowstone National Park, Wyoming, USA. January 2016
Grizzly bear (Ursus arctos horribilis) female with sub-adult cub rolling on back, in grassland. Yellowstone National Park, Wyoming, USA. June
Sunrise over Madison Mountains, Beaverhead National Forest, Montana, USA, July 2011
RF - Red fox (Vulpes vulpes) foraging in snow covered valley. Hayden Valley, Yellowstone National Park, USA. February 2019. (This image may be licensed either as rights managed or royalty free.)
Red fox (Vulpes vulpes) sequence of snow diving whilst hunting for rodents. Hayden Valley, Yellowstone National Park, USA. February 2019. Digital composite
North American river otter (Lutra canadiensis), two standing at edge of frozen Upper Yellowstone River, Hayden Valley, Yellowstone, USA. February
Steam rising from Grand Prismatic thermal pool in snow covered landscape. Midway Geyser Basin, Yellowstone National Park, USA. February 2019
Coyote (Canis latrans) standing near waters edge, in snow. Hayden Valley, Yellowstone National Park, USA. January 2019
Grizzly bear (Ursus arctos) running through water, Lake Clark National Park, Alaska, September
Soaptree yucca (Yucca elata) and twilight sky, showing Belt of Venus (blue of earth's shadow on the atsmophere)Soaptree yucca (Yucca elata) and twilight sky, showing Belt of Venus (blue of earth's shadow on the atsmophere below pink of sunset). White Sands National Monument, New Mexico, USA. November
Orange bush monkeyflower (Mimulus aurantiacus). Native to California and Oregon, USA. Focus stacked
Yellow mariposa lily (Calochortus luteus) with dark nectar guides. Cultivated in glasshouse, Surrey, England, UK. Endemic to California
Tulip tree (Liriodendron tulipifera) flower with orange band to attract bees, produces copious nectar. In visible light. Surrey, England, UK. Native to USA. Focus stacked. Sequence 1/2
Yellow Labrador retriever in fresh snow, Clinton, Connecticut, USA
Great egret (Ardea alba) with prey in beak, in evening light. Myakka River State Park, Florida, USA, March. Sequence 2/2
Great egret (Ardea alba) diving head into water to catch prey, in evening light. Myakka River State Park, Florida, USA, March. Sequence 1/2
Tricoloured heron (Egretta tricolor) fishing by flying low over water, amongst Water lilies. Everglades National Park, Florida, USA. March
