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Pere Davids deer / Milu (Elaphurus davidianus) during the rutting season, stag, Hubei Tian ezhou Milu National Nature Reserve, Shishou, Hubei, China
Impala (Aepyceros melampus) allo-grooming, iMfolozi game reserve, KwaZulu-Natal, South Africa, August
Red fox (Vulpes vulpes) vixen grooming cub, Sado Estuary, Portugal. May
Egyptian goose (Alopochen aegyptiaca) pair amongst Devil s-thorn yellow flowers (Tribulus terrestris) Etosha National Park, Namibia, March
Head portrait of black Andalusian stallion and mare meeting for the first time in southern Spain, Europe
Jaguar (Panthera onca) mother grooming four month cub, native to Southern and Central America, captive
Grey wolf (Canis lupus) mother and two month old cubs, pups begging for food by licking mothers mouth, captive
Bornean orangutan (Pongo pygmaeus) mother and babies, Tanjung Puting National Park, Borneo-Kalimatan, Indonesia, endangered species
Erect-crested penguin (Eudyptes sclateri) pair in greeting display. Antipodes Island, New Zealand Sub-Antarctic Islands. Endemic to Antipodes and Bounty Islands. Endangered species
RF- Gentoo Penguin (Pygoscelis papua) chick begging parent for food, Antarctica
Emperor Penguin aka (Aptenodytes forsteri) adult penguins with their chick, Weddell Sea, Gould Bay, Antarctica
Flightless cormorant (Phalacrocorax harrisi) pair in courtship Puerto Pajas, Isabela Island, Galapagos
Tiger (Panthera tigris), female and cub, Ranthambhore National Park, Rajasthan, India
Tabby kittens, Stanley and Fosset, 3 months old, sitting and snuggling together
Lionesses (Panthera leo) grooming each other, Masai-Mara Game Reserve, Kenya
Baby Grey Squirrel (Sciurus carolinensis) kissing a tabby kitten
West African / Niger Giraffe (Giraffa camelopardalis peralta) mother and baby. Zoo of Doue La Fontaine, France. Captive
Grey heron (Ardea cinerea) and Great egret (Ardea alba) fighting over fish, Lake Csaj, Pusztaszer, Hungary, December. Winner of the Portfolio category of the Terre Sauvage Nature Images Awards
Tabby kittens, Stanley and Fosset, 12 weeks old, walking together
African lions (Panthera leo) one lioness licking another, Okavango Delta, Botswana
A wild newborn colt (Equus caballus) standing up being nuzzled by its mother. Foothills of Reno, Nevada, USA, May
The Lovers Baobabs (Adansonia grandidieri). Near Morondava, Madagascar" The Lovers" Baobabs (Adansonia grandidieri). Near Morondava, Madagascar, Africa, September 2010
Two Cheetahs (Acinonyx jubatus) touching noses in greeting display, Serengeti NP, Tanzania, East Africa, January
Mating pair of King Penguins (Aptenodytes patagonicus) on Gold Beach, South Georgia Island, Sub Antarctica
RF- Gannets (Morus) displaying, Great Saltee, County Wexford, Ireland. May
RF - Head portrait of black Andalusian mare and stallion meeting for the first time in Spain, Europe
Tabby kittens, Picasso and Smudge, with paws on each others shoulders
Gannet (Morus bassanus) breeding pair during part of their elaborate courtship ritual. Saltee Islands, Republic of Ireland, May
African Lion (Panthera leo) cub reaches for a moment of intimacy with its father, Northern Tuli Game Reserve, Botswana
Wild Chincoteague (Equus caballus) two breeding stallions greeting one another, Chincoteague National Wildlife Refuge, Chincoteague Island, Virginia, USA, June
Waved albatross (Phoebastria irrorata) courting pair mutually grooming. Punta Cevallos, Espanola (Hood) Island, Galapagos, Ecuador, May
Giraffe (Giraffa camelopardalis) female bending down to calf, Okavango Delta, Botswana
Black-tailed Prairie Dogs (Cynomys ludovicianus), two pups (right) seem to be kissing their parent. The behaviour is a gesture of recognition
Young sandy rabbits kissing
Silky sifaka (Propithecus candidus) pair in tree, Marojejy National Park, Madagascar, Endangered species
African lion (Panthera leo) lioness licking cub, Sabi Sand Game Reserve, South Africa, June
Female Polar bear with very small cubs {Ursus maritimus} Canada
Two Cheetah (Acinonyx jubatus) brothers grooming one another. bonding behaviour, Linyanti, Chobe National Park, Botswana
Grey seal (Halichoerus grypus) pup and mother sniffing and nuzzling each other. Donna Nook, Lincolnshire, UK. November
Two Miniature American shepherd puppies, aged 7 weeks, sitting side by side, portrait
Yellow Labrador retriever puppy and Ragdoll-cross kitten side by side, touching noses, portrait
Long-eared owls (Asio otus) autumn, two owls interacting while roosting in tree, social behaviour, The Netherlands
Two Konik wild horses (Equus ferus caballus) foals nuzzling, Meinerswijk nature reserve, near Arnhem, the Netherlands. June
Two Lionesses (Panthera leo) nuzzling surrounded by three cubs, Kgalagadi Transfrontier Park, Northern Cape, South Africa
Lowland gorilla (Gorilla gorilla) mother and young in forest, Loango National Park, Gabon. January
Eurasian lynx (Lynx lynx) kitten, aged six weeks, showing affection towards its mother. Breeding and reintroduction program, Germany. Captive. July
RF - Eurasian lynx (Lynx lynx) kittens, aged eight months, showing affection towards each other, breeding and reintroduction program, Germany. Captive
