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Bornean orangutan (Pongo pygmaeus) mother and babies, Tanjung Puting National Park, Borneo-Kalimatan, Indonesia, endangered species
Erect-crested penguin (Eudyptes sclateri) pair in greeting display. Antipodes Island, New Zealand Sub-Antarctic Islands. Endemic to Antipodes and Bounty Islands. Endangered species
RF- Gentoo Penguin (Pygoscelis papua) chick begging parent for food, Antarctica
Emperor Penguin aka (Aptenodytes forsteri) adult penguins with their chick, Weddell Sea, Gould Bay, Antarctica
Flightless cormorant (Phalacrocorax harrisi) pair in courtship Puerto Pajas, Isabela Island, Galapagos
Tiger (Panthera tigris), female and cub, Ranthambhore National Park, Rajasthan, India
Lionesses (Panthera leo) grooming each other, Masai-Mara Game Reserve, Kenya
Baby Grey Squirrel (Sciurus carolinensis) kissing a tabby kitten
West African / Niger Giraffe (Giraffa camelopardalis peralta) mother and baby. Zoo of Doue La Fontaine, France. Captive
Grey wolf (Canis lupus) mother and two month old cubs, pups begging for food by licking mothers mouth, captive
Grey heron (Ardea cinerea) and Great egret (Ardea alba) fighting over fish, Lake Csaj, Pusztaszer, Hungary, December. Winner of the Portfolio category of the Terre Sauvage Nature Images Awards
Tabby kittens, Stanley and Fosset, 12 weeks old, walking together
African lions (Panthera leo) one lioness licking another, Okavango Delta, Botswana
A wild newborn colt (Equus caballus) standing up being nuzzled by its mother. Foothills of Reno, Nevada, USA, May
Domestic pig, free range sow and piglet sniffing noses, UK, August 2010
A pair of Australian Sea Lions (Neophoca cinerea) socialising in the shallows. Kangaroo Island, South Australia, Australia. Southern Ocean. November
Two Cheetahs (Acinonyx jubatus) touching noses in greeting display, Serengeti NP, Tanzania, East Africa, January
Roan antelope (Hippotragus equinus) with young offspring, Mlilwane nature reserve breeding programme, Swaziland, Southern Africa
Two Cape Ground squirrels (Xerus inauris) greeting, Kgalagadi Transfrontier Park, South Africa Non-ex
RF- Gannets (Morus) displaying, Great Saltee, County Wexford, Ireland. May
RF - Head portrait of black Andalusian mare and stallion meeting for the first time in Spain, Europe
Tabby kittens, Picasso and Smudge, with paws on each others shoulders
Gannet (Morus bassanus) breeding pair during part of their elaborate courtship ritual. Saltee Islands, Republic of Ireland, May
African Lion (Panthera leo) cub reaches for a moment of intimacy with its father, Northern Tuli Game Reserve, Botswana
Giraffe (Giraffa camelopardalis) female bending down to calf, Okavango Delta, Botswana
Unravelling Fern fronds (Unknown species). Near Nepenthes Field Camp, mid-altitude montane forest. Centre of Maliau Basin, Sabahs Lost World, Borneo
Two newly fledged burrowing owl chicks (Athene cunicularia) one being groomed by its mother (far left) Pantanal, Brazil. WINNER
Two Cheetah (Acinonyx jubatus) brothers grooming one another. bonding behaviour, Linyanti, Chobe National Park, Botswana
Grey seal (Halichoerus grypus) pup and mother sniffing and nuzzling each other. Donna Nook, Lincolnshire, UK. November
Two Miniature American shepherd puppies, aged 7 weeks, sitting side by side, portrait
Yellow Labrador retriever puppy and Ragdoll-cross kitten side by side, touching noses, portrait
Long-eared owls (Asio otus) autumn, two owls interacting while roosting in tree, social behaviour, The Netherlands
Pair of European bee-eaters (Merops apiaster) with courtship offering of insect prey, Pusztaszer, Kiskunsagi National Park, Hungary
Two Konik wild horses (Equus ferus caballus) foals nuzzling, Meinerswijk nature reserve, near Arnhem, the Netherlands. June
Two Lionesses (Panthera leo) nuzzling surrounded by three cubs, Kgalagadi Transfrontier Park, Northern Cape, South Africa
Lowland gorilla (Gorilla gorilla) mother and young in forest, Loango National Park, Gabon. January
Eurasian lynx (Lynx lynx) kitten, aged six weeks, showing affection towards its mother. Breeding and reintroduction program, Germany. Captive. July
RF - Eurasian lynx (Lynx lynx) kittens, aged eight months, showing affection towards each other, breeding and reintroduction program, Germany. Captive
Wild Welsh pony colt greeting his father, Carneddau Mountains, Snowdonia, Wales, UK. June
Black rhino (Diceros bicornis) mother and calf, Solio Game Reserve, Laikipia, Kenya. September
Impala (Aepyceros melampus) with redbilled oxpecker (Buphagus erythrorhynchus), Kruger National Park, South Africa
Fairy Tern (Gygis alba) pair, Cousin Island, Republic of Seychelles
Leopard (Panthera pardus) female grooming cub age 6 months Little Kwara, Botswana June
Bornean orangutan (Pongo pygmaeus) mother and baby, Tanjung Puting National Park, Borneo-Kalimatan, Indonesia, endangered species
Cheetah (Acinonyx jubatus) two leaning close together, Okavango Delta, Botswana
Coastal brown bear (Ursus arctos) and cub, Lake Clarke National Park, Alaska, September
Wild Mustang pinto foal nuzzling up to mother, Sand Wash Basin Herd Area, Colorado, USA
Wild Mustang foals among wild flowers, Pryor Mountains, Montana, USA
