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Atlantic salmon (Salmo salar) portrait, migrating to spawn in river, Gaspe Peninsula, Quebec, Canada, October
Three spot damselfish (Dascyllus trimaculatus) with Sea anemone home, Yap, Micronesia
Common anemonefish (Amphiprion perideraion) with eggs in Magnificent sea anemone (Heteractis magnifica) Yap, Micronesia
Striped mackerel (Rastrelliger kanagurta) with mouths wide open as they swim through the water, filtering zoopankton with their gill rakers. Marsa Shouna, Port Ghalib, Marsa Alam, Egypt. Red Sea
Bluestreak cleaner wrasse (Labroides dimidiatus) sticking its head into the mouth of a Red Sea fusilier (Caesio lunaris) during a cleaning session. Small Crack, Sha ab Mahmud, Sinai, Egypt. Red Sea
Coral Rabbitfish (Siganus corallinus) portrait, Tubbataha Reef Natural Park, UNESCO World Heritage Site, Sulu Sea, Cagayancillo, Palawan, Philippines
Cichlids (Cichlidae) in Lake Malawi, Malawi, November. Photographed for The Freshwater Project
Striped mackerel (Rastrelliger kanagurta) shoal swimming with open mouths, filtering for zooplankton. Marsa Shouna, Port Ghalib, Marsa Alam, Egypt. Red Sea
Bluespine unicornfish (Naso unicornis) Shark Reef to Jolande, Ras Mohammed National Park, Egypt, Red Sea
Trumpetfish (Aulostomus maculatus) using a Spanish Hogfish (Bodianus rufus) as a stalking horse. Spanish Hogfish is eating an arm of a starfish. Bonaire, Leeward Antilles, Caribbean
Oceanic whitetip shark (Carcharhinus longimanus) accompanied by pilotfish (Naucrates ductor) as it cruises beneath the surface of the Red Sea in the late afternoon, close to Little Brother Island
Puffin (Fratercula arctica) standing on Sea thrift (Armeria maritima) with wings out stretched with a beak full of sand eels. Close-up portrait. Fair Island, Shetland Islands, Scotland, July
Portrait of Blacktip grouper (Epinephelus fasciatus) resting in Barrel sponge (Xestospongia testudinaria) Mabul Island, Sabah, Borneo, Malaysia. Sulawesi Sea
Oceanic whitetip shark (Carcharhinus longimanus) accompanied by a group of Pilotfish (Naucrates ductor). Rocky Island, Egypt. Red Sea
School of large Atlantic bluefin tuna (Thunnus thynnus) captive in growing pen. Depsite looking much smaller in the photo, these fish were 1.5m (5ft) long. Malta. Mediterranean Sea
The vertical reef wall at Shark Reef, Ras Mohammed, with Scalefin anthias (Pseudanthias squamipinnis) and soft corals (Dendronephthya spp.) Yellowsaddle goatfish (Parupeneus cyclostomus)
Large male Napoleon wrasse (Cheilinus undulatus) yawning. Ras Mohammed, Sinai, Egypt. Red Sea
School of Longnose parrotfish (Hipposcarus harid) swarming over reef. Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
RF- School of Blackfin barracuda (Sphyraena qenie) forming circle in open water at Shark Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
Klunzingers wrasse (Thalassoma klunzingeri) swimming over the edge of fringing reef where the water depth drops instantly from few feet to hundreds of meters. Ras Mohammed Marine Park, Sinai, Egypt
Male Yellow-headed jawfish (Opistognathus aurifrons) blows out the clutch of eggs he is incubating in his mouth to oxygenate them. East End, Grand Cayman, Cayman Islands, British West Indies
A female (in front) and male Sockeye salmon (Oncorhynchus nerka) over eggs in their spawning river. Adams River, British Columbia, Canada, October
Pair of Sockeye salmon (Oncorhynchus nerka) on their redd in a shallow stream. Female in front of male. Huihil Creek, Adams River, British Colombia, Canada, October
Striped mackerel (Rastrelliger kanagurta) mouth wide open as it swims through the water, filtering zoopankton with its gill rakers. Marsa Shouna, Port Ghalib, Marsa Alam, Egypt. Red Sea
RF - A vibrant Red Sea coral reef scene, with orange female Scalefin anthias fish (Pseudanthias squamipinnis) teeming over Fire coral (Millepora dichotoma)
RF - Close up detail of the eye and skin pattern of a Giant hawkfish (Cirrhitus rivulatus). Socorro Island, Pacific Ocean
RF - Banggai cardinalfish (Pterapogon kauderni). Lembeh Strait, North Sulawesi, Indonesia
Striped marlin (Kajikia audax) feeding on bait balls, Eastern Pacific, Thetis Bank, Baja California, Mexico
Blue and gold fusilier / Scissor tailed fusilier (Caesio caerulaurea) school swimming through shafts of sunlight, West Papua, Indonesia
RF - School of large Atlantic bluefin tuna (Thunnus thynnus) captive in a growing pen. Depsite looking much smaller in the photo, these fish were 1.5m (5ft) long. Malta. Mediterranean Sea
Thousand of Gilt-head bream (Sparus aurata) inside a sea cage used for aquaculture, Ponza Island, Italy, Tyrrhenian Sea, Mediterranean
Ruffe (Gymnocephalus cernua) shoal, Danube Delta, Romania, June
Mandarinfish (Synchiropus splendidus) pair spawning. The larger male (behind) is lifting the female into the spawning rise on his pectoral fin, the smaller female (foreground) is releasing eggs
Blue spotted trevally (Caranx bucculentus) hunting small fish in the reef, South Ari Atoll, Maldives, Indian Ocean
Red Sea anemonefish (Amphiprion bicinctus) in their home, a Sea anemone (Heteractis magnifica), on a coral reef. Big Brother island, Red Sea. Egypt
Atlantic Puffin (Fratercula arctica) in flight, coming into land with beak full of sand eels (Ammodytes tobianus). Fair Isle, Shetland Islands, Scotland, UK, July
Three False clownfish (Amphiprion ocellaris) in Sea anemone (Heteractis magnifica) Lighthouse Reef, Cabilao Island, Bohol, Central Visayas, Philippines, Pacific Ocean
Atlantic salmon (Salmo salar) migrating for spawning in river, Gaspe Peninsula, Quebec, Canada, October
Oceanic manta ray (Mobula birostris) accompanied by three Black jacks (Caranx lugubris) silhouetted, Revillagigedo Islands, Mexico, Pacific Ocean
Detail of the yellow and blue stripes of an adult Emperor angelfish (Pomacanthus imperator) on a coral reef. Laamu Atoll, Maldives. Indian Ocean
Large whip goby (Bryaninops amplus) with a parasitic copepod (with two trailing egg ribbons) perched on Red whip coral. Whale Rock. Fiabacet Islands, Misool, Raja Ampat, West Papua, Indonesia
Three Sockeye salmon/ Red salmon (Oncorhynchus nerka) swimming upstream as they migrate back to the river of their birth to spawn
Tail-spot blenny (Ecsenius stigmatura) peering out from amongst the coral, Triton Bay, West Papua, Indonesia, Pacific Ocean
Female scalefin anthias (Pseudanthias squamipinnis) swimming in front of a school of glassfish (Apogonidae), whose shiny bodies are rendered into out of focus bokeh patterns. Laamu Atoll, Maldives
School of powderblue surgeonfish (Acanthurus leucosternon) feeding, Laamu Atoll, Maldives. Indian Ocean
Kelp bass (Paralabrax clathratus) in Giant kelp (Macrocystis pyrifera) forest, Santa Barbara, California, USA
RF - Portrait of a coral grouper (Cephalopholis miniata) next to a sea fan (Annella sp.) on a coral reef. Raja Ampat, West Papua, Indonesia. Ceram Sea
Slate pencil sea urchins (Heterocentrotus mammillatus) on coral reef with Pyramid butterflyfish (Hemitaurichthys polylepis), Hawaii
