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RF - Arctic Fox (Vulpes lagopus) portrait in winter coat, Svalbard, Norway, April
Blue sky reflected in ripples of water, Svalbard, Norway, June 2010
Summer Forget-me-not (Anchusa azurea) in flower, Chania, Crete, April
Corys shearwaters (Calonectris diomedea) diving among a mass of shoaling fish to feed, along with Atlantic spotted dolphins (Stenella frontalis), Formigas Islet dive site, Azores, Portugal
Blue whale (Balaenoptera musculus) underwater, Coronado Islands, Baja California, Mexico, Eastern Pacific Ocean
Giant Manta Ray (Manta birostris), San Benedicto Island, Revillagigedo Archipelago Biosphere Reserve (Socorro Islands), Pacific Ocean, Western Mexico. Vulnerable species
Adelie penguin (Pygoscelis adeliae) jumping from ice edge into the sea, Antarctica
Blue lupin (Lupinus angustifolius) Sierra de Grazalema Natural Park, southern Spain, May
Blacktip reef shark (Carcharhinus melanopterus) juvenile in shallow water, Maldives, Indian Ocean
RF - School of large Atlantic bluefin tuna (Thunnus thynnus) captive in a growing pen. Depsite looking much smaller in the photo, these fish were 1.5m (5ft) long. Malta. Mediterranean Sea
Orcas / killer whales (Orcinus orca) swimming in open water, Three Kings Islands, New Zealand. Pacific Ocean
Blue iridescent scales of a Nymphalid butterfly (Epiphile orea) magnified 11x, deceased insect found in Piedade, Sao Paulo, Brazil. South-east Atlantic Forest
Striped marlin (Kajikia audax) feeding on bait balls, Eastern Pacific, Thetis Bank, Baja California, Mexico
Close-up of Hyacinth Macaw (Anodorhynchus hyacinthinus feathers, Brazil
DELETE Striped marlin (Kajikia audax) feeding on bait balls, Eastern Pacific, Thetis Bank, Baja California, Mexico
Yellow springtails (Symphypleona) on Forget-me-not flower (Myosotis) Zillertal Alps, Austria, July
Swift (Apus apus) flock screaming in flight against blue sky and clouds, Monmouthshire, Wales, UK, July
Patters of sunlight on a shallow sandy seabed. North Sound, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea
Brandts cormorant (Phalacrocorax penicillatus) bursts through a school of Pacific chub mackerel (Scomber japonicus) while hunting beneath an oil rig
Forget-me-not flowers (Myosotis arvensis) on woodland floor, Norfolk, England, UK, March
A split level image of Southern stingray (Hypanus americanus) swimming over a sand bar. Grand Cayman, Cayman Islands. Caribbean Sea. Digital composite
Adelie penguin (Pygoscelis adeliae) diving near ice flow, Antarctica. Small reproduction only
Stars and Milky Way over Durdle Door and the Jurassic Coast, with the lights of Weymouth and Portland beyond, Dorset, England, UK, December 2013
Blue triggerfish (Pseudobalistes fuscus) Egypt, Red Sea
RF- Small jellyfish (Pelagia noctiluca) swimming beneath surface. Marine Protected Area of Portofino (Area Marina Protetta, Portofino), Liguria, Italy. Mediterranean Sea
Lago di Landro at dawn, Dolomite Mountains, Sud Tirol / Alto Adige, ItalyLago di Landro at dawn, Dolomite Mountains, Sud Tirol/Alto Adige, Italy
Long-beaked common dolphins (Delphinus capensis) feeding in Sardines, (Sardinops sagax) with snorkeller behind, Eastern Cape, South Africa
Peacock (Pavo cristatus feather close up showing iridescence at 10x magnification
Southern stingray (Dasyatis americana) swimming over sand in shallow water, split level photo with blue sky and clouds. The Sandbar, Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea
School of Blackfin barracuda (Sphyraena qenie), Kimbe Bay, West New Britain, Papua New Guinea
Close-up of a Jays (Garrulus glandarius) wing, showing blue covert feathers
Brick wall in the Blue City, Jodhpur, Rajasthan, India
The Blue City, Jodhpur, Rajasthan, India. March 2015
Common blue butterfly (Polyommatus icarus) female, Southwest London, England, UK. June
Morpho butterfly (Morpho peleides) photographed on white background. Costa Rica, March
Yellow and blueback fusilier (Caesio teres) schooling over coral reef, West Papua, Indonesia
Blue and gold fusilier / Scissor tailed fusilier (Caesio caerulaurea) school swimming through shafts of sunlight, West Papua, Indonesia
Ice cave beneath the Breidemerkurjokull glacier, eastern Iceland, March 2015
Selfoss in winter, Vatnajokull National Park, Northeast Iceland, March 2016
Godafoss in winter, Bardardalur district of North-Central Iceland, March 2016
Ice in the glacial lagoon at Jokulsarlon, Iceland, February 2015
Ice cave below the Breidamerkurjokull Glacier, eastern Iceland, February 2015
Landscape of Hvannadalshnukur, Iceland, February 2015
Details of ice in an ice cave below the Breidamerkurjokull Glacier, eastern Iceland, February
RF - Young lionfish (Pterois volitans) swimming near surface hunting silversides, at dusk. Beacon Rock, Sha ab Mahmood, Sinai, Egypt. Red Sea
RF - Titan triggerfish (Balistoides viridescens) swimming over hard coral (Acropora sp.) gardens. Kri Island, Raja Ampat, West Papua, Indonesia. Dampier Strait, tropical west Pacific Ocean
RF - Groyne at Dawlish Warren, Devon, UK. January 2016
RF- Elongate surgeonfish (Acanthurus mata) school. Indonesia
Wren (Troglodytes troglodytes) on frost covered fern, Dunham Massey, Cheshire, UK. Winner of the Habitat Category of the British Wildlife Photographer of the Year Awards (BWPA) 2017
Small tortoiseshell butterfly (Aglais urticae) resting on old painted door. Dorset, UK, August
Napoleon wrasse (Cheilinus undulatus) portrait. Red Sea, Egypt. October
Sunbeams below the surface of the sea, Guadalupe Island, Baja California, Mexico. East Pacific Ocean
Yellow and blueback fusilier (Caesio teres) shoal, Similan Islands, Andaman Sea, Thailand
Trumpet / Stemless Gentian flower (Gentiana acaulis) looking down the middle of the flower, showing five-fold symmetry. Nordtirol, Austrian Alps. June
Hyacinth macaws (Anodorhynchus hyacinthinus) mutual preening, Estrada Parque, Pantanal, Brazil
Cirrus clouds displaying wind shear, Brechin, Scotland, UK, September
Loch Beinn a Meadhoin in the mist, Inverness, Scotland, December 2013
Young Polar bear (Ursus maritimus) walking across melting sea ice, Scott Inlet, Baffin Island, Canadian Arctic, August. Vulnerable species
Polar bear (Ursus maritimus) mother and large second year cub swim across calm water, Scott Inlet, Baffin Island, Canadian Arctic, August. Vulnerable species
Common magpies (Pica pica) perched on frost covered branches, Jvaskyla, Finland, January
Schooling Scalloped hammerhead sharks (Sphyrna lewini) Cocos Island National Park, Costa Rica
Assumption of Mary Pilgrimage Church, in twilight, reflected in Lake Bled, Bled, Slovenia, October 2014
Bigeye trevally or jack (Caranx sexfasciatus) shoal, Sipadan, Malaysia
Emperor penguin (Aptenodytes forsteri), Amanda Bay, Prydz Bay, Ingrid Christensen Coast, East Antarctica, November
Snow petrel (Pagodroma nivea) flying, Antarctica, November
Lesser flamingo (Phoeniconaias minor) flock, aerial view, Magadi lake, Kenya, September
Sandy beach and bright blue ocean, Fuerteventura, Canary Islands. April 2013
Cracked mud and blue stones, Fuerteventura, Canary Islands
Blue sea star (Linckia laevigata) on coral reef, Fiji, South Pacific
Broadclub cuttlefish (Sepia latimanus) Komodo National Park, Indonesia
Dawn over the mountains and Nam Song River at Vang Vieng, Laos, March 2009
Stone on rock pattern, Kilve beach, Somerset, UK. January 2014
Atlantic bluefin tuna (Thunnus thynnus) within tuna farm, containing around 1000 per net. Saint Pauls Bay, Malta. Mediterranean Sea
Fulmar (Fulmarus glacialis) flying over sea, Iceland
Close up of pattern of adult Emperor angelfish (Pomacanthus imperator) Tulamben, Bali, Indonesia. Java Sea
Detail of pattern of an adult Emperor angelfish (Pomacanthus imperator) Seraya, Tulamben, Bali, Indonesia. Java Sea
Large school of Bohar snappers (Lutjanus bohar) gather with smaller shoal of Batfish (Platax orbicularis) in open water off Ras Mohammed at the tip of Sinai, Egypt
School of Bohar snappers (Lutjanus bohar) Shark Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
Male Bullethead parrotfish (Chlorurus sordidus) displaying in late afternoon. Ras Katy, Sinai, Egypt. Gulf of Aqaba, Red Sea
Male Scalefin anthias (Pseudanthias squamipinnis) swimming in open water, Ras Katy, Sharm El Sheikh, Sinai, Egypt. Gulf of Aqaba, Red Sea