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Two paint horses, a palomino and a sorrel quarter horse running, Flitner Ranch, Shell, Wyoming, USA
Four chocolate working cocker spaniels on wall. Mother and offspring. Monmouth, Monmouthshire, Wales
Cavalier King Charles Spaniel puppies aged 7 weeks, with tricolour and blenheim colouration, on hay bale in straw
A shiver of Caribbean reef sharks (Carcharhinus perezi) swim over a coral reef with Common sea fans (Gorgonia ventalina) and sea plumes (Pseudopterogorgia sp)
Japanese cranes (Grus japonensis) displaying in snow, Hokkiado, Japan, February
Peacock butterfly (Inachis io) and Small tortoiseshell butterflies (Aglais urticae) feeding on Buddleia flowers (Buddleia davidii), Derbyshire, UK, September
Brown hare, (Lepus europaeus), group of animals in field, Islay, Scotland, UK. March
Four Cedar waxwings (Bombycilla cedrorum) feeding on Crabapple (Malus sp) fruit, Ithaca, New York, USA, February
White ibis (Eudocimus albus) group of four in flight above clouds, Fort Myers Beach, Gulf Coast, Florida, USA, March
Silvery grebe (Podiceps occipitalis) attempting to join Hooded grebes (Podiceps gallardoi) during a breeding display, Patagonia National Park, Santa Cruz, Argentina. Cropped
Great crested grebe (Podiceps cristatus) carrying chicks on its back in early morning light, Valkenhorst Nature Reserve, The Netherlands, Europe. May
Asiatic lion (Panthera leo persica), two females and two cubs. Gir National Park, Gujarat, India. Photo© Phillip Ross/Felis Images
Wood Ducks (Aix sponsa), females behave aggressively toward a male (second from right) that is trying to join them on a floating log, autumn, New York, USA, October
Four black-and-white Border Collie puppies, age 7 weeks, sitting in a row
Trumpeter swans (Cygnus buccinator) on the edge of the Upper Yellowstone River. Hayden Valley, Yellowstone National Park, Wyoming, USA. January
Pairs of Scarlet Macaws (Ara macao) in flight. Osa Peninsula (near Corcovado National Park), Costa Rica, Central America
Group of four Giraffes (Giraffa camelopardalis) Mkuze, South Africa
Warthogs (Phacochoerus aethiopicus) mother with babies drinking, Mkhuze game reserve, KwaZuiu-Natal, South Africa, June
Moutain gorilla (Gorilla beringei beringei) close up of hand, Virunga National Park, North Kivu, Democratic Republic of Congo, Africa, Critically endangered
Horsesgrazing on Bulbarrow Hill at dawn, Dorset, England, UK, July 2014
RF - Eschmeyers scorpionfish (Rhinopias eschmeyeri) composite image showing different colour variations on black background. Lembeh Strait, North Sulawesi, Indonesia
Giant Otters (Pteronura brasiliensis) running along a sand bar. Cuiaba River, Pantanal, Brazil
Galah (Cacatua roseicapilla) in tree, Piccaninny Plains. Cape York Peninsula, Queensland, Australia
Rocky mountain bighorn sheep (Ovis canadensis canadensis) grazing. Lamar Valley, Yellowstone National Park, Wyoming, USA. January
Yacare Caiman (Caiman yacare) gaping to regulate its body temperature, with attendant hoverflies, the Paraguay River, Taiama Reserve, western Pantanal, Brazil, South America
Atlantic walruses (Odobenus rosmarus rosmarus) hanging out in shallow water in Svalbard, , Norway, June
RF - King penguin (Aptenodytes patagonicus), four returning to sea. St Andrews Bay, South Georgia. November. (This image may be licensed either as rights managed or royalty free.)
Eastern white-bearded wildebeest (Connochaetes taurinus), four running in line, on migration, Masai Mara National Reserve, Kenya
Common rue (Ruta graveolens) flower in visible light, nectar at base of petals. Stamens flip up sequentially with one still folded over the receptacle. Surrey, England, UK
Common rue (Ruta graveolens) flower in UV light, pollen and nectar at base of petals fluorescing. Insect pollinators are able to see position of rewards on offer. Surrey, England, UK
Scarlet Macaws (Ara macao) drinking water inside an Amazonian claylick. Yasuni National Park, Orellana, Ecuador
Silhouette of Red deer (Cervus elaphus) stag, hind and fawns at sunset, Norfolk, UK
Gannet (Morus bassanus) adults and a youngster silhouetted against an orange red sky as another adult prepares to land. Shetland Islands, Scotland, UK. September
Hazel nuts (Corylus avellana) Dorset, UK September
Sika deer (Cervus nippon) stags in summer coat, Arne RSPB Reserve, Dorset, England, August
Black-bellied whistling-ducks (Dendrocygna autumnalis), group perched amid Spanish moss on bald cypress tree, Lakeland, Florida, USA, January
Chilean devil rays / Mobulas (Mobula tarapacana) offshore Santa Maria, Azores, Portugal, Atlantic Ocean
Domestic piglets {Sus scrofa domestica} Devon, UK
Common Spotted Orchid {Dactylorhiza fuchsii} flowers, Hardington Moor NNR, Somerset, England, June
Four Common / Plains Zebras (Equus quagga) standing side by side, portrait, Okavango Delta, Botswana, Africa
Three Brown noddies / Common noddies (Anous stolidus) trying to steal a fish from a Brown pelican (Pelecanus occidentalis), Santiago Island, Galapagos Islands, Pacific Ocean
Female Cheetah (Acinonyx jubatus) with juveniles crossing grassland in rain storm, Kgalagadi Transfrontier Park, Northern Cape, South Africa
Four White storks (Ciconia ciconia) perched on street lights, silhouetted at dusk, Madrid, Spain. June
Four Grey partridges (Perdix perdix) on snow-covered arable field, Scottish Borders, Scotland, UK. January
Four Wolves (Canis lupus) walking in snow, Yellowstone National Park, USA. January
RF - Group of four Mycena toadstools on mossy stump. The New Forest, Hampshire, UK. October. (This image may be licensed either as rights managed or royalty free)
Three female Putorana snow sheep (Ovis nivicola borealis) climbing up rocky mountainside with lamb following, Putoransky State Nature Reserve, Putorana Plateau, Siberia, Russia
Four Hereford bullocks looking over a fence with Cow Parsley (Anthriscus sylvestris) Worcestershire, England, May
Fiordland penguins (Eudyptes pachyrhynchus) walk along a forest path, heading back to their burrows, after foraging in the ocean. North of Hst, Westland District, South Island, New Zealand
Red stingray (Hemitrygon akajei), four in a row, viewed from below. Chiba Prefecture, Honshu, Japan. October
Himalayan griffon vulture (Gyps himalayensis) Angsai Nature Reserve (Valley of the Cats), Sanjiangyuan National Nature Reserve, Tibetan Plateau, Qinghai, China
Group of Longfin batfish (Platax teira) beneath the surface, close to an island. Farondi Islands, Misool, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean
Coyote (Canis latrans) foraging in deep winter snow disturbed by grazing American bison (Bison bison) Hayden Valley, Yellowstone National Park, Wyoming, USA. January
Four Pygmy seahorses (Hippocampus bargibanti) on a Seafan / Gorgonian (Muricella sp. ) including one or several pregnant males. New Caledonia, Pacific Ocean
RF - Cape buffalo (Syncerus caffer) drinking at night, Zimanga private game reserve, South Africa. (This image may be licensed either as rights managed or royalty free.)
Bar headed geese (Anser indicus), four in flight, one calling. Coming in to land at breeding site, Qinghai Lake, Qinghai Province, China
Mountain hare, (Lepus timidus), four animals on snowy hillside in winter, Scotland, UK.January
Mountain hare, (Lepus timidus), four animals on snowy hillside in winter, Scotland, UK, February
Sika deer (Cervus nippon) amongst flowering heather, Arne RSPB reserve, Dorset, England, UK, August
Fledgling Barn swallows (Hirundo rustica) calling for food in barn. Dorset, UK, August
RF - Herring gull (Larus argentatus) with blue sky, England, UK, Digital composite
Andean flamingos (Phoenicopterus andinus), Laguna Colorada, Bolivia. December 2016
Florida manatee (Trichechus manatus latirostris), four resting on sandy river bed. Split level image. Three Sisters Spring, Crystal River, Florida, USA
Griffon vulture (Gyps fulvus) landing amongst other vultures on cliff above sea, Liendo Valley within the Montana Oriental Costera, Cantabria, Spain
Family of Whooper swan (Cygnus cygnus), Iceland, February
Brown bear (Ursus arctos) mother leads three cubs over hillside in snow, Kamchatka, Far east Russia, May
Haflinger horse mares and foals running in meadow. England, UK
Feral domestic rabbits (Oryctolagus cuniculus) sitting up alert, Okunojima Island, also known as Rabbit Island, Hiroshima, Japan
Svalbard reindeer (Rangifer tarandus platyrhynchus) mother and calves, Svalbard, Norway, February
Horses hoof / Tinder fungus (Fomes fomentarius) on tree trunk, European beech (Fagus sylvatica) forest, Stuzica primeval Forest, Unesco World Heritage Site, Poloniny National Park
Mallard (Anas platyrhynchos) ducklings, Bradfield Nature Reserve, Berkshire, England, UK, May
Meerkats (Suricata suricatta) standing alert, Kgalagadi Transfrontier Park, Northern Cape, South Africa, January
Alpaca Herd on Norfolk Farm, winter
Grey-cheeked parakeets (Brotogeris pyrrhoptera) perched and grooming on a branch. Guayaquil, Guayas, Ecuador
RF - Four colourful Red-and-green macaws or Green-winged macaws (Ara chloropterus) in flight over forest canopy. Mato Grosso do Sul, Brazil. September
Barn owl (Tyto alba) four chicks in nest, La Pampa, Argentina
Four California spiny lobsters (Panulirus interruptus) shelter beneath a boulder in a kelp forest. Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America
Bluebells (Endymion non-scriptus) flowering, Perthshire, Scotland, May
Cape buffalo (Syncerus caffer) crossing the Chobe River, followed by swarm of flies, Chobe Game Reserve, Botswana, Africa
Spotted eagle ray (Aetobatus narinari) school, Kagi kandhu, ; North Male Atoll, Maldives, Indian Ocean
Cheetah (Acinonyx jubatus) cubs 4 months, Masai-Mara Game Reserve, Kenya. Vulnerable species
Blacktip reef sharks (Carcharhinus melanopterus) swimming in shallow crystal clear water, Aldabra Atoll, Seychelles, Indian Ocean
Four Wisent / Bison (Bison bonasus) standing in a row. Bialowieza Forest, Bialowieza National Park, Poland, November