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Shoebill stork (Balaeniceps rex) in the swamps of Mabamba, Lake Victoria, Uganda
Purple toadflax (Linaria purpurea), on verge of footpath, South Croydon, Surrey, England, June
Spotted lanternfly (Lycoma delicatula) fourth instar nymph, Wissahickon Valley Park, Pennsylvania, USA, July
Axolotl (Ambystoma mexicanum), white or leucistic form, neotenic salamander. Captive, critically endangered in the wild, occurs in Mexico
Sea elephant (Cardiopoda placenta), marine pelagic gastropod mollusc in the family Pterotracheidae. Deep sea heteropod. Atlantic Ocean, close to Cape Verde. Controlled conditions
Northern pike (Esox lucius) amongst Shining pondweed (Potamogeton lucens). Lake Neuchatel, Switzerland, September. Photographed for The Freshwater Project
Shoebill stork flying (Balaeniceps rex) in the swamps of Mabamba, Lake Victoria, Uganda
Shoebill stork (Balaeniceps rex) eating a fish in the swamps of Mabamba, Lake Victoria, Uganda
Osprey (Pandion haliaetus) landing at its nest in a Bald cypress tree (Taxodium distichum) draped with epiphytic Spanish moss (Tillandsia usneoides). Blue Cypress Lake, Florida, USA. April
Hanging thief robber fly (Diogmites sp) Philadelphia, Pennsylvania, USA, July
Pacific white-sided dolphins (Lagenorhynchus obliquidens) underwater, Queen Charlotte Strait, British Columbia, Canada. September
Giant Pacific octopus (Enteroctopus dofleini) showing arms and suckers, Vernon Rock, Queen Charlotte Strait, British Columbia, Canada. September
Roe deer (Capreolus capreolus) buck with well developed horns standing on garden lawn carpeted with Common daisies (Bellis perennis) and Buttercups (Ranunculus acris) near flowering Lupins (Lupinus)
Portrait of a female rescued Grey-headed flying-fox (Pteropus poliocephalus) pup. Captive - Taken in controlled conditions and under supervision of wildlife carer
Roe deer (Capreolus capreolus) buck running past a garden shed at night, Wiltshire, UK, April. Property released
European badger (Meles meles) walking down garden steps at night, with house in background, Wiltshire, UK, June
European badger (Meles meles) juvenile walking down some garden steps at night, flanked by flowering Helebores (Helleborus sp.) Catmint (Nepeta sp.) and Geraniums, Wiltshire, UK, June
Young Roe deer (Capreolus capreolus) buck with developing horns in velvet approaching a garden pond, Wiltshire garden, UK, February
European badger (Meles meles) reflected in a garden pond as it drinks from it at night, Wiltshire, UK, June. Property released
Alert young Roe deer (Capreolus capreolus) buck with developing horns standing on a terraced garden lawn, Wiltshire garden, UK, October
Painted lady butterfly (Vanessa cardui) feeding on knapweed Sutcliffe Park Nature Reserve, Eltham, London, England, UK. June
Common meadow-rue (Thalictrum flavum), locally rare plant, Chertsey Meads, Surrey, England, June
Autumn Lady s-tresses orchid (Spiranthes spiralis) locally rare plant. Epsom Downs, Surrey, England, August
Common poppies (Papaver rhoeas) in field, Chicklade, Wiltshire, England, July
Marbled white butterfly (Melanargia galathea) on Wild marjoram (Origanum vulgare) Farthing Downs, Surrey, England, July
Pale toadflax (Linaria repens), a rare plant in Surrey. Park Downs, Chipstead Downs SSSI, Surrey, England, July
Yellow water-lily (Nuphar lutea) Lake Morat, Canton Vaud, Switzerland, July. Photographed for The Freshwater Project
Inser Weiher in autumn, a pond with red coloured water, caused by dissolved tannins. Berner Seeland, Berne, Switzerland, November. Photographed for The Freshwater Project
Yellow water-lily (Nuphar lutea) underwater. Lake Morat, Vaud, Switzerland, September. Photographed for The Freshwater Project
Bewicks swan (Cygnus columbianus bewickii) adults swimming on a marshland pool, Gloucestershire, UK, November
Dense flock of Common teal (Anas crecca) with a few Wigeon (Anas penelope) and Northern shoveler (Anas clypeata) in flight over others resting on partly frozen flooded marshes
Common shelduck pair (Tadorna tadorna) standing and drinking in the margins of a shallow lake, Gloucestershire, UK, November
Mute swan (Cygnus olor) pair preening in unison, Gloucestershire, UK, December
Marsh harrier (Circus aeruginosus) female hunting among a panicked flock of Common teal (Anas crecca) in flight over marshland, RSPB Greylake Nature Reserve, Somerset, UK, December
Hairy violet (Viola hirta) clump flowering in a chalk grassland meadow, Wiltshire, UK, March
Green-winged orchid (Orchis / Anacamptis morio) clump, pale pink / white form, Mellis Common, Suffolk, UK, May
Germander speedwell (Veronica chamaedrys) clump flowering in a chalk grassland meadow, Wiltshire, UK, May
Portrait of male silverback Mountain gorilla (Gorilla beringei beringei) feeding. Bwindi Impenetrable Forest National Park, Uganda, Africa
Great blue turaco (Corythaeola cristata) in Bigodi wetland santuary, Uganda
Olive baboon (Papio hamadryas anubis) mother with babies in a tree. Kibale National Park, Uganda, Africa
Chimpanzee (Pan troglodytes schweinfurthii) male vocalising in tree, Kibale National Park, Uganda, Africa
Mountain gorilla (Gorilla beringei) mother with one month baby feeding in tree. Member of Katwe group. Bwindi Impenetrable Forest National Park, Uganda, Africa
Army ant (Eciton burchellii) carrying prey, Copalinga Reserve, Ecuador
Long-wattled umbrellabird (Cephalopterus penduliger) Buenaventura Reserve, Ecuador
Vochysia tree and rainforest landscape, Copalinga Reserve, Ecuador, December 2018
Goitered or Black-tailed gazelle (Gazella subgutturosa) male, Kalamaili National Nature Reserve, Xinjiang, China
Siberian ibex (Capra sibirica) silhouetted on ridge, Tian Shan mountains, Xinjiang, China
Bactrian camel (Camelus bactrianus) male silhouetted at sunset, living in the wild but owned by a camel herdsman, Turpan Basin, Gobi Desert, Xinjiang, China
Steller sea lions (Eumetopias jubatus) playing underwater, Inian Island, Alaska, USA. August
Split level of Sunflower stars (Pycnopodia helianthoides) on rocks underwater, Minstrel Island, British Columbia, Canada. March
Split level of Acorn barnacles (Balanus glandula) just below the surface, Seven Tree Island, Browning Pass, Queen Charlotte Strait, British Columbia, Canada. September
Black-and-white sea flea (Chromopleustes oculatus) on a Sunflower star (Pycnopodia helianthoides), Port Neville, Johnstone Strait, British Columbia, Canada. March
Split-level of Cormorants (Phalacrocorax sp. ) standing on rocks with Sea lettuce (Ulva spSplit-level of Cormorants (Phalacrocorax sp.) standing on rocks with Sea lettuce (Ulva sp.) seen underwater, Hussar Bay, Nigei Island, Queen Charlotte Strait, British Columbia, Canada. September
Fried egg jellyfish (Phacellophora camtschatica) viewed from below; Browning Pass, Queen Charlotte Strait, British Columbia, Canada. September
A Giant rock scallop (Crassadoma gigantea) encrusted with and surrounded by several other invertebrates animals, including White anemones (Metridium sp.), White mushroom ascidians (Distaplia sp.)
Gooseneck barnacles (Pollicipes Polymerus) amongst kelp, Nakwakto Rapids, Slingsby Channel, British Columbia, Canada. September
Harbor seal (Phoca vitulina) using a large, encrusted boulder as a tool to scratch itself against, Gowland Harbour, Quadra Island, British Columbia, Canada. March
Split level of a Lions mane Jellyfish (Cyanea capillata) and Feather-boa kelp (Egregia menziesii), Seven-tree Island, Browning Pass, Queen Charlotte Strait, British Columbia, Canada. September
Split level of Sockeye salmon (Oncorhynchus nerka) during migration; Adams River, British Columbia, Canada. October
Moon jellyfish (Aurelia labiata) viewed from below, Browning Pass, Queen Charlotte Strait, British Columbia, Canada. February
Thousands of Pacific Sandlance (Ammodytes hexapterus) swim overhead, as a few rockfish (Sebastes sp.) swim in the distance
Moon jellyfish (Aurelia labiata) viewed from below; Agamemnon Channel, Sunshine Coast, British Columbia, Canada. March
Purple and Ochre sea stars (Pisaster ochraceus) preying on Pacific blue mussels (Mytilus trossulus)
Sockeye salmon (Oncorhynchus nerka) migration, Adams River, British Columbia, Canada. October
Sockeye salmon (Oncorhynchus nerka) males fighting in river Adams River, British Columbia, Canada. October
Bull kelp canopy (Nereocystis luetkeana) floating on the surface providing protective habitat for juvenile Rockfishes (Sebastes spp.) Browning Pass, Queen Charlotte Strait, British Columbia, Canada
Northern basket star (Gorgonocephalus eucnemis) Slingsby Channel, British Columbia, Canada. May
Sea fleas (Chromopleustes oculatus) on dorsal surface of a Giant sunflower seastar, Port Neville, Johnstone Strait, British Columbia, Canada. March
Proliferating anemones (Epiactis prolifera) and Kelp lace bryozoans (Membranipora serrilamella) on Bull kelp (Nereocystis luetkeana), Browning Pass, Queen Charlotte Strait, British Columbia, Canada
Wolf eel (Anarrhichthys ocellatus) Croker Rock, Queen Charlotte Strait, British Columbia, Canada. September
Stubby squid (Rossia pacifica) Nigei Island, Queen Charlotte Strait, British Columbia, Canada
Split level shot of kelp and other seaweeds partly exposed at low tide, with coniferous forest in the background
Schooling rockfishes (Sebastes spp. ) with Bull kelp (Nereocystis luetkeana)Schooling rockfishes (Sebastes spp.) with Bull kelp (Nereocystis luetkeana). Different species of rockfish often swim together in mixed schools, as here
Early morning view, looking southeast from Gordon Channel. Gordon Channel, Queen Charlotte Strait, British Columbia, Canada. September
Opalescent nudibranchs (Hermissenda crassicornis) and Mushroom compound ascidians (Distaplia occidentalis). Browning Pass, Queen Charlotte Strait, British Columbia, Canada. September
Purple-ringed Topsnail (Calliostoma annulatum), Orange Social Tunicate (Metandrocarpa taylori) and Lattice-work Bryozoan (Phidolophora pacifica), Browning Pass, Queen Charlotte Strait
Opalescent nudibranch (Hermissenda crassicornis) preys upon stinging hydroids. Browning Pass, Queen Charlotte Strait, British Columbia, Canada. September
Northern kelp crab (Pugettia producta) large adult male is clinging to strands of bull kelp (Nereocystis luetkeana). Nigei Island, Queen Charlotte Strait, British Columbia. September
Lions Mane Jellyfish (Cyanea capillata) with Blue rockfish (Sebastes mystinus) stealing food from the jellyfishs tentacles. Hunt Rock, Queen Charlotte Strait, British Columbia, Canada. September
Moon jelly (Aurelia labiata) and Cross jellies (Mitrocoma cellularia) split level view in Browning Pass, Queen Charlotte Strait, British Columbia. October
Hooded nudibranchs (Melibe leonina) with oral hood for catching planktonic prey. In the Pacific Northwest they are often found clinging to bull kelp (Nereocystis luetkeana)
Maroon hermit crabs (Pagurus hemphilli) in shells of Blue Topsnail (Calliostoma ligatum) Browning Pass, Queen Charlotte Strait, British Columbia, Canada. September
Helmet crab (Telmessus cheiragonus) and Hooded nudibranchs (Melibe leonina), Nigei Island, Queen Charlotte Strait, British Columbia, Canada. October
Harbor seal (Phoca vitulina) near the Wreck of the Themis, Croker Rock, Queen Charlotte Strait, British Columbia. September