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Adult Mountain gorilla (Gorilla beringei beringei) holding baby, Hirwa group, Volcanoes National Park, Rwanda
Osprey (Pandion haliaetus) with Atlantic needlefish (Strongylura marina) it has caught. Blue Cypress Lake, Florida, USA, April
Hybrid Poplar tree plantation in autumn (Populs sp. ). Boardman Tree Farm, near IrrigonHybrid Poplar tree plantation in autumn (Populs sp.). Boardman Tree Farm, near Irrigon, Oregon, USA. October
Common loon (Gavia immer) with a chick alongside. British Columbia, Canada. June
Common eider duck (Somateria mollissima) female defending her nest from arctic fox (Vulpes lagopus). Longyearbyen, Spitsbergen. June
American lobster (Homarus americana) in eelgrass (Zostera marina). Nova Scotia, Canada. July
Stellers jay (Cyanocitta stelleri) perched on branch. British Columbia, Canada, June
European green crab (Carcinus maenas), an invasive species in North America, Kejimkujik Seaside National Park, Nova Scotia, Canada. July
Pelagic thresher shark (Alopias pelagicus) being cleaned by a bluestreak cleaner wrasse (Labroides dimidiatus) at Monad Shoal off Malapascua Island, Philippines
Larval Wonderpus octopus (Wunderpus photogenicus) drifting in the open ocean at night off Anilao, Philippines
Wonderpus octopus (Wunderpus photogenicus) in its juvenile or larval stage. At this stage it is transparent and drifts in the open ocean as plankton. Anilao, Philippines
Coastal taipan (Oxyuranus scutellatus) tasting air, Julatten, north Queensland, Australia
Pine tree (Pinus sp) in winter, Bryce Canyon National Park, Utah, USA, January
Cheviot sheep grazing eroding machair at front of sand dunes, North Uist, Scotland, UK, June
Salt cones on the Salar de Uyuni, Bolivia. March. The Salar is the worlds largest salt flat, at over 10500 square kilometres. Salt is shoveled into these cones, to be collected later. March 2012
Chestnut-eared aracari (Pteroglossus castanotis) on cat-gut vine. Pantanal, Brazil, August
Common grackle pair (Quiscalus quiscula) in courtship display. Wakodahatchee Wetlands, Florida, USA, April
Snowy egret (Egretta thula) shows its breeding plumage. St. Johns Management Area, Florida, USA, March
Quiver trees (Aloidendron dichotomum) silhouetted against dawn sky, Namibia, May
Yellow warbler (Setophaga petechia aureola) on lava rocks. Isabela Island, Galapagos. May
Great frigatebird (Phaethon aethereus) male at nest site, with gular sac inflated to attract females. Genovesa Island, Galapagos, May
Violet-tailed sylph hummingbird (Aglaiocercus coelestis) in flight, about to land. Ecuador, May
Blue tit (Cyanistes caeruleus) bringing Crab spider (Misumena vatia) prey to nest, Monmouthshire Wales, UK, May
Scottish wildcat (Felis silvestris grampia) captive, endemic to Scotland. September
Blue shark (Prionace glauca) with a pair of pilot fish (Naucrates ductor) off Halifax, Nova Scotia, Canada. July
Sugar kelp (Sacchinaria latissima) off Nova Scotia, Canada. September
A female American black bear (Ursus americanus) and her young cub, Nova Scotia, Canada. July
Head of a North Atlantic right whale (Eubalaena glacialis) showing callosities, pathches of roughened skin that are unique to each whale. Gulf of Saint Lawrence, Canada. IUCN Status: Endangered
Aerial view of Carters Beach on the Southshore of Nova Scotia, Canada. August
Bottlenose dolphin (Tursiops truncatus) underwater playing in the shallows off Eleuthera, Bahamas
Moorish idol (Zanclus cornutus) juvenile with transparent skin in the open ocean at night off Anilao, Philippines
Queen conch (Lobatus gigas) juvenile feeding on algae growing on Seagrass (Thalassia testudinum) blades. Eleuthera, Bahamas
Ribbon eel ( Rhinomuraena quaesita) Lembeh Strait, North Sulawesi, Indonesia
Nudibranch (Chromodoris willani) with a scale worm. Lembeh Strait, North Sulawesi, Indonesia
Corkscrew or Long tentacle anemone (Macrodactyla doreensis). Lembeh Strait, North Sulawesi, Indonesia
Boxer Crab (Lybia tessellata) with sea anemones in its claws for defense. Lembeh Strait, North Sulawesi, Indonesia
Taylors pygmy leatherjacket (Brachaluteres taylori). Lembeh Strait, North Sulawesi, Indonesia
Deckens sifaka (Propithecus deckenii) grooming each other, Tsimembo area, Madagascar. Lenses for Conservation project
Madame Berthes Mouse Lemur (Microcebus berthae), the worlds smallest primate, Kirindy forest, Madagascar. Lenses for Conservation project
Blue morning flower (Ipomoea indica) flowers and carpet of leaves. Cyprus. April
White asphodel (Asphodelus albus) flowerhead. Cyprus. April
Ashy mining bee (Andrena cineraria), feeding on Dandelion (Taraxacum offinicale) Monmouthshire, Wales, UK. September
Captive reared juvenile White stork (Ciconia ciconia) in flight over the Knepp Estate soon after release, Sussex, UK, August 2019
Wild cat (Felis sylvestris) portrait, part of captive breeding project at Alladale Estate, Scotland, UK, October
Purple vipers bugloss (Echium plantaineum) growing in fields of Corn Marigold (Chrysanthemum)Purple vipers bugloss (Echium plantaineum) growing in fields of Corn Marigold (Chrysanthemum segetum) Project Neptune National Trust, Cornwall, England, UK, June
Lesser horseshoe bats (Rhinolophus hipposideros) in magnesium mine, Shropshire, England, UK, April
European Beaver (Castor fiber) amongst Pondweed (Potomogeton natans) in freshwater lake Bevis Trust, Carmarthenshire, Wales, UK, June
Sequence of strips of cultivated Machair with early shoots of Black oats (Avena strigosa) pushing through the sand. North Uist, Scotland, UK, June
Corncrake (Crex crex) walking at dawn on Machair, North Uist, Scotland, UK, June
Dunlin (Calidris alpina) small wader with young in machair. North Uist, Scotland, UK, June
Two Little terns (Sterna albifrons) at the nest amongst Black oats (Avena strigosa) growing on shell rich sands Machair, North Uist, Scotland, UK, June
Little tern (Sterna albifrons) at the nest amongst Black oats (Avena strigosa) growing on shell rich sands Machair, North Uist, Scotland, UK, June
Little tern (Sterna albifrons) with newly hatched chick, Gronant dunes, Denbighshire, Wales, UK, June
Little tern (Sterna albifrons ) feeding sand eel (Hyperoplus spp) to young chick. Gronant Dunes, Denbighshire, Wales, UK, June
Strips of cultivated Black oats (Avena strigosa), grown by crofters, who create sillage and feed it to their cattle in the winter. Berneray, North Uist, Scotland, UK, July
Little tern (Sterna albifrons) nest with three eggs in shingle ridge. County Wicklow, Ireland, June
Corn Marigold (Chrysanthemum segetum) and Scentless mayweed (Tripleurospermum inodorum) machair habitat, North Uist, Scotland, UK, July
Red Deer (Cervus elaphus) portrait of head of stag and antlers. Alladale Estate, Scotland, UK, October
Dunlin (Calidris alpina ) North Uist, Outer Hebrides, Scotland, UK, June
Red deer (Cervus elaphus) on moorland / blanket bog silhouetted against rushes, Alladale Estate, Scotland, UK, October
Shoreline of Lake Powell, seen from Alstrom Point on Romana Mesa. Glen Canyon National Recreation Area, on the Arizona/Utah border, USA, January
Brown bear Ursus arctos) searching for clams on tidal mudflat with waterbirds. Lake Clark National Park, Alaska. August
Hoodoo formation with caprock, at Stud Horse Point, Utah, January
James flamingo / Puna flamingo (Phoenicoparrus jamesi). Lago Colorado, Bolivia. March
King penguins (Aptenodytes patagonicus) walking in line on a windy beach. Sanders Island, Falklands. November
Yacare caiman (Caiman yacare) Cuiaba River, Pantanal, Brazil, August
Southern red-billed hornbill male (Tockus rufirostris) Mahatsu Reserve, Botswana, June
Common giraffe (Giraffa tippelskirchi) bending to drink at a waterhole, with another standing on the opposite side of the pool reflected in the water
Purple gallinule (Porphyrio martinicus) reaches for giant bulrush (Schoenoplectis californicus) seeds. Wakodahatchee Wetlands, Florida, USA, April
Roseate spoonbill (Platalea ajaja) at its nesting colony site, in early morning light. St. Johns Management Area, Florida, USA, April
Late afternoon light warms the colors in the Yellow Mounds area, Badlands National Park, South Dakota, USA, September
Black-browed albatross pair (Thalassarche melanophris) mutual grooming during copulation. New Island, Falkland Islands, November
Large leaves on Pacific blackberry (Rubus ursinus) above bracken fern and horsetail, is a small wet area. Oregon, USA. June
Namaqua chameleon (Chamaeleo namaquensis) catching larvae with its extended tongue. In the Namib desert near Swakopmund, Namibia
Inside an ice cave in the Vatnajokull Glacier, near Jokulsarlon, Iceland, March
Snow monkey or Japanese macaque (Macaca fuscata) female with baby clinging to her back. Jigokudani, Japan, February
Sally lightfoot crabs (Grapsus grapsus) on the beach at Puerto Egas, Santiago Island, Galapagos, May
Western screech owl (Megascops kennicottii) is camouflaged as it looks from a tree cavity. Bosque del Apache National Wildlife Refuge, New Mexico, USA, January
Soaptree yucca (Yucca elata) at sunset. White Sands National Monument, New Mexico, USA. January
Cheetah (Acinonyx jubatus) stretches on a downed tree. Zimbabwe. September
Yellow legged mining bee (Andrena flavipes) female at nest burrow, River Monnow, Monmouthshire, Wales, UK, April
Mining bee (Andrena sp. ), harvesting pollen from anthers of Cherry tree (Prunus spMining bee (Andrena sp.), harvesting pollen from anthers of Cherry tree (Prunus sp.) Ripon Wisconsin USA, May
Painted nomad bee (Nomada fucata) male bee with characteristic green eyes, sleeping with jaws locked around grass stem, Monmouthshire, Wales, UK. June
Smeathmans furrow bee (Lasioglossum smeathmanellum), feeding on wild alpine strawberry (Fragaria vesca), Monmouthshire, Wales, UK. December