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American bison (Bison bison) female covered in hoar frost near hot spring, portrait. Midway Geyser Basin, Yellowstone National Park, USA. February
Puma (Puma concolor puma) young male standing with mouth open. Estancia Amarga, near Torres del Paine National Park, Patagonia, Chile. December
Orca (Orcinus orca) hunting South American sealion (Otaria flavescens) close to shore. Punta Norte, Valdez Peninsula, Argentina. April
Red squirrel (Sciurus vulgaris), two curled up asleep in drey inside nest box. Nest of lichen and Pine needles. Highlands, Scotland, UK. Medium repro only
Blue poppy (Meconopsis zangnanensis) Mt Qomolangma National Park, Qinghai Tibet Plateau, China
Red fox (Vulpes vulpes) in mid air, snow diving / pouncing whilst hunting for rodents. Hayden Valley, Yellowstone National Park, USA. February
Sumatran pit viper (Trimeresurus sumatranus) juvenile on tree trunk, in tropical rainforest. Danum Valley, Sabah, Borneo, Malaysia
Flying fish (Exocoetidae) in Sargasso Sea, Atlantic Ocean
Dugong (Dugong dugon) male and juvenile Golden trevally (Gnathanodon speciosus) swimming over Seagrass (Halophila stipulacea) meadow. Marsa Nabaa, Marsa Alam, Egypt
Siberian weasel (Mustela sibirica) standing in snow. Jiudingshan Nature Reserve, Mao Country, Sichuan Province, China. November
Wild Jaguar (Panthera onca), Endangered, Cuiaba River, Pantanal, Mato Grosso, Brazil
Giant river otter (Pteronura brasiliensis) resting on fallen tree trunk above Cuiaba River. Pantanal, Mato Grosso, Brazil
Little red flying fox (Pteropus scapulatus) female and suckling baby in flight, searching for place to roost. Nitmiluk National Park, Northern Territory, Australia
Cape buffalo (Syncerus caffer caffer) portrait, Red-billed oxpeckers (Buphagus erythrorhynchus) removing parasites. Nakuru, Kenya
Spanish lavender (Lavandula stoechas). Faia Brava Reserve, Archaeological Park of the Coa Valley, Western Iberia, Portugal. April
Orca (Orcinus orca) with South American sealion (Otaria flavescens) in mouth, beaching itself on shore. Punta Norte, Valdez Peninsula, Argentina. April
Reef manta ray (Manta alfredi) shoal filter feeding on plankton concentrated by monsoon currents in Raa Atoll, Vandhoo Thila, Maldives. October
Lenticular clouds above Torres del Paine National Park. Patagonia, Chile. November 2018
Guanaco (Lama guanicoe) female, alert whilst looking at Puma (Puma concolor) on on opposite slope, portrait. Torres del Paine National Park, Patagonia, Chile. November
Tussah moth (Antheraea godmani) close-up of head and bipectinate antennae. Santa Clara, Chiriqui Province, Panama. Focus stacked image
White rhino (Ceratotherium simum) female and calf looking in opposite directions, reflected in waterhole at night. Zimanga private game reserve, KwaZulu-Natal, South Africa
Johnston's three-horned chameleonJohnston's three-horned chameleon, (Trioceros johnstoni), male on tree branch, Kahuzi-Biega NP, Democratic Republic of Congo, November. Non-ex
Round-leaved sundew (Drosera rotundifolia), close up. Bystock Pools, Devon Wildlife Trust nature reserve, near Exmouth, Devon. June. Focus stacked image
Western diamondback rattlesnake (Crotalus atrox) camouflaged amongst rocks, with rattle raised. Sonoran Desert, Arizona, USA. Controlled conditions. June 2019
Turtle dove (Streptopelia turtur) female feeding chicks on nest. Ranmore Common, near Dorking Surrey, England, UK. July
Moss-covered Sitka spruce (Picea sitchensis) and sword fern (Polystichum munitum) in the Hoh Rain Forest, Olympic National Park, Washington, USA, June
Kestrel (Falco tinnunculus) male hovering overhead, Cornwall, UK, April
Canada Goose (Branta canadensis) in flight. Gilbert Water Ranch, Arizona, USA. January
Okapi (Okapia johnstoni). Beauval Zoo Parc, France. Captive
Melanistic leopard / Black panther (Panthera pardus fusca) resting in tree. Nagarhole National Park, India. Photo© Phillip Ross/Felis Images
Fishing cat (Prionailurus viverrinus) walking with fish in mouth. Andhra Pradesh, India. Camera trap image. Photo© Anjani Kumar/Felis Images
Asiatic lion (Panthera leo persica), two females and two cubs. Gir National Park, Gujarat, India. Photo© Phillip Ross/Felis Images
Golden langur (Trachypithecus geei) sitting in tree, Umanada Temple Island, Assam, India
Eastern grey kangaroo (Macropus giganteus), two males fighting. Grampians National Park, Victoria, Australia
Female cheetah (Acinonyx jubatus) named Silgi (means bright future in Swahili) walking with 7 cubs. Masai Mara National Reserve, Kenya
Magnificent frigatebird (Fregata magnificens) in flight over sea between Isabela Island and Fernandina Island, Galapagos
Galapagos sea lion (Zalophus wollebaeki) jumping out of Pacific ocean. Sombrero Chino, Santiago Island, Galapagos
Alcedo giant tortoise (Chelonoidis vandenburghi), group bathing. Alcedo Volcano, Isabela Island, Galapagos. 2017
RF - Red fox (Vulpes vulpes) foraging in snow covered valley. Hayden Valley, Yellowstone National Park, USA. February 2019. (This image may be licensed either as rights managed or royalty free.)
RF - Puma (Puma concolor puma), young male walking on hillside near Torres del Paine National Park, Patagonia, Chile. November. (This image may be licensed either as rights managed or royalty free.)
RF - King penguin (Aptenodytes patagonicus), four returning to sea. St Andrews Bay, South Georgia. November. (This image may be licensed either as rights managed or royalty free.)
RF - Guanaco (Lama guanicoe) standing in front of mountain towers of Paine. Torres del Paine National Park, Patagonia, Chile. December 2018
Southern elephant seal (Mirounga leonina), two males, equally matched with mouths open in aggression. Out of focus King penguin (Aptenodytes patagonicus) colony in background
Towers and Central Massif reflected in Lago Pehoe at sunrise. Torres del Paine National Park, Patagonia, Chile. November 2018
Red fox (Vulpes vulpes) sequence of snow diving whilst hunting for rodents. Hayden Valley, Yellowstone National Park, USA. February 2019. Digital composite
Rafflesia (Rafflesia keithii) flower aged approximately 3 days on rainforest floor. Lower slopes of Mt Kinabalu, Sabah, Borneo, Malaysia
Black-browed albatross (Thalassarche melanophris) in flight over sea at sunrise. South Atlantic Ocean between The Falklands and South Georgia. November
North American river otter (Lutra canadiensis), two standing at edge of frozen Upper Yellowstone River, Hayden Valley, Yellowstone, USA. February
Darwins rhea (Pterocnemia pennata), two with tower peaks in background. Torres del Paine National Park, Patagonia, Chile. December
Guanacos (Lama guanicoe), five silhouetted on slope in evening. Torres del Paine National Park, Patagonia, Chile. December
Steam rising from Grand Prismatic thermal pool in snow covered landscape. Midway Geyser Basin, Yellowstone National Park, USA. February 2019
American bison (Bison bison) female grazing near thermal pool, snow on ground. Biscuit Geyser Basin, Yellowstone National Park, USA. February 2019
Burnt-tip orchid (Neotinea ustulata) group flowering in species rich ancient alpine meadow. North Tyrol, Austria. June
American bison (Bison bison), two females covered in hoar frost near hot spring, portrait. Midway Geyser Basin, Yellowstone National Park, USA. February
Coyote (Canis latrans) standing near waters edge, in snow. Hayden Valley, Yellowstone National Park, USA. January 2019
Morning Glory thermal pool surrounded by snow, coniferous forest in background. Near Daisy Geyser Basin, Yellowstone National Park, USA. January 2019
Common knapweed (Centaurea nigra) in ancient alpine meadow with hut. in distance, North Tyrol, Austria. June 2018
Fragrant orchids (Gymnadenia conopsea) amongst wildflowers in ancient alpine meadow. North Tyrol, Austria. June
Guanaco (Lama guanicoe) standing, towers of Torres del Paine National Park in background. Patagonia, Chile. December 2018
King penguin (Aptenodytes patagonicus), two sleeping on snow. Grytviken, South Georgia. March
Central Massif and towers of Torres del Paine National Park at sunrise, island reflected in Lago Pehoe in foreground. Patagonia, Chile. November 2018
Grizzly bear (Ursus arctos) running through water, Lake Clark National Park, Alaska, September
Mute swan (Cygnus olor), two cygnets sheltering under parent's wingMute swan (Cygnus olor), two cygnets sheltering under parent's wing. Richmond Park, London, England, UK. April
Mute swan (Cygnus olor) swimming with cygnets in misty lake in morning light. Richmond Park, London, England, UK. May. Cropped image
Red deer (Cervus elaphus) stag bellowing during rut in grassland, doe in background
Southern stingray (Dasyatis americana) two swimming over sand bar, under stormy sky. Grand Cayman, Cayman Islands. British West Indies
Spinner dolphin (Stenella longirostris) pod socialising close to surface. Fury Shoal, Sataya Reef, Marsa Alam, Egypt
Oceanic whitetip shark (Carcharhinus longimanus) swimming close to surface. Brothers Islands, Egypt
Hawfinch 1+Coccothraustes coccothraustes+2 perched on branch amongst Hawthorn 1+Crataegus sp+2 berries. Sierra de Grazalema Natural Park, Cadiz, Spain. February
Supermoon rising over Stonehenge, the biggest for 68 years. Wiltshire, England, UK. November 2016
Mandarin duck (Aix galericulata) reflected in water. Richmond Park, London, England, UK
Mute swan (Cygnus olor) cygnet swimming, cocking head. Barnes, London, England, UK. April
Small tortoiseshell (Aglais urticae) butterfly flying amongst Scabious (Knautia arvensis) in hay meadow. Wensum Valley, Norfolk, England, UK. July
American quarter horse standing in forest, portrait. Alberta, Canada. February
European mantis (Mantis religiosa) hunting swarming Pale burrower mayfly (Ephoron virgo), mayflies recently hatched. Tudela, La Ribera de Navarra, Navarre, Spain. August
American quarter horse, three standing in snow. Alberta, Canada. February
Southern royal albatross (Diomedea epomophora) sub-adults, group displaying in tussock grassland. Campbell Island, New Zealand. February
Chinese goral (Naemorhedus griseus) in snow. Jiudingshan Nature Reserve, Mao Country, Sichuan Province, China. November
Snow partridge (Lerwa lerwa) flock flying past snowy rockface, in evening light. Jiudingshan Nature Reserve, Mao Country, Sichuan Province, China. November
Coscoroba swan (Coscoroba coscoroba) at nest with eggs. La Pampa, Patagonia, Argentina. February
Greater rhea (Rhea americana) nest with many eggs, at edge of arable field. Patagonia, Argentina. October
Hyacinth macaw (Anodorhynchus hyacinthinus) pair perched on branch, silhouetted under crescent moon. Mato Grosso Forest, Pantanal, Mato Grosso, Brazil
Marsh deer (Blastocerus dichotomus) walking across track in forest. Pantanal, Mato Grosso, Brazil
Cheetah (Acinonyx jubatus) walking, silhouetted at dusk. Zimanga private game reserve, KwaZulu-Natal, South Africa
