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Light shining into Abismo Anhumas or Anhumas Abyss. This is a 80 metre deep lake, at the bottom of a 72 metre deep cave
Under water in the Abismo Anhumas or Anhumas Abyss. This is a 80 metre deep lake, at the bottom of a 72 metre deep cave
Split level view of blue green waters of Verzasca River flowing over granite rocks at Lavertezzo, Canton Tessin, Switzerland, September. Taken for the Freshwater Project
RF - Lions (Panthera leo) - two brothers patrolling territorial boundary. Short grass plains on the border of Serengeti / Ngorongoro Conservation Area (NCA) near Ndutu, Tanzania
Eastern chipmunk (Tamias striatus) gathering chestnuts, Chestnut Hill, Philadelphia, Pennsylvania, USA, September
Jocotoco antpitta (Grallaria ridgelyi) Tapichalaca Biological Reserve, Ecuador
Yellow-plumed parakeets (Leptosittaca branickii) at nest cavity in wax palm stump, Tapichalaca Biological Reserve, Ecuador
Starling (Sturnus vulgaris) feeding fledgling chicks in urban garden. Greater Manchester, UK. May
Salt marsh cord grass (Spartina alterniflora) on shore of Cape Cod, with Long Point Lighthouse in the background, Massachusetts, USA, September
Cat gecko (Aeluroscalabotes felinus) with coiled tail, Sarawak, Malaysian Borneo
Sumatran pit viper (Trimeresurus sumatranus) female in rain forest understorey. Danum Valley, Sabah, Borneo
Bottlenose dolphin (Tursiops truncatus) at surface, Moray Firth, Highlands, Scotland. May
RF - Starling (Sturnus vulgaris) feeding fledgling chicks in urban garden. Greater Manchester, UK. May
Fire salamander (Salamandra salamandra) female almost ready to give birth to her larvae into the stream, Apennines. Antola Regional Park, Italy
RF - Arctic fox (Vulpes lagopus) male, blue colour morph. Hornstrandir Nature Reserve, Iceland. March
Rangimairewhenua or Blue Lake, Nelson Lakes National Park, Southern Alps, New Zealand, February 2013. Photographed for The Freshwater Project
Huntsman spider (Megaloremmius leo) Andasibe, Madagascar. Endemic species
Lance-nosed chameleon (Calumma gallus), Andasibe-Mantadia National Park. Madagascar
Dried up river bed in the Anti Atlas mountains of Morocco, North Africa. April 2012. In recent years, rainfall totals have reduced by around 75% as a result of climate change
Great Fen / Raft spider (Dolomedes plantarius), adult female eating an invasive species of fish, Western mosquitofish, (Gambusia affinis), Alessandria, Italy, August
Lions (Panthera leo) - two brothers patrolling territorial boundary, affectionate behaviour, border of Serengeti / Ngorongoro Conservation Area (NCA) near Ndutu, Tanzania
Snapping turtle (Chelydra serpentina) with Painted turtle (Chrysemys picta) feeding on algae on the back of the snapper, Maryland, USA. August
RF -Three colourful Red-and-green macaws (Ara chloropterus) in flight over forest canopy. Mato Grosso do Sul, Brazil. September
RF - Four colourful Red-and-green macaws or Green-winged macaws (Ara chloropterus) in flight over forest canopy. Mato Grosso do Sul, Brazil. September
Wavy-lined emerald moth caterpillar (Synchlora aerata) which camouflages itself by attaching parts of flower to itself, on Black-eyed susan (Rudbeckia hirta) Morris Arboretum, Philadelphia
Passionflower (Passiflora cubalensis) Yanacocha Reserve, Ecuador
RF - Bottlenose dolphins (Tursiops truncatus) porpoising, Chanonry Point, Moray Firth, Highlands, Scotland. August
Canada geese (Branta canadensis) flock in flight, Moray Firth, Highlands, Scotland. June
Horned adder (Bitis caudalis) waiting for a passing prey into the Namib Desert, Brandberg, Namibia
Crack of the continental drift between the North American and Eurasian Plates at Thingvellir, Thingvellir National Park, UNESCO World Heritage Site, Iceland. September 2009
Lesser kestrel (Falco naumanni) male flying past during courtship with Mole cricket in its talons. Extremadura, Spain
Clown frog (Dendropsophus leucophyllatus), Villa Carmen Biological Station, Peru
Horned adder (Bitis caudalis) close up of eye, Brandberg area, Namibia
Two male Lesser kestrels (Falco naumanni) in pursuit flight, one driving off the other from its nesting territory. Extremadura, Spain
Laguna Miluni, contaminated with mine effluent and with a low level due to drought. This reservoir is fed by glacial meltwater from the Andean peak, Huayna Potosi, Bolivian Andes. October 2015
Black and white image of English oak tree (Quercus robur) with long exposure of wind blown clouds, Monmouthshire, Wales, UK, September 2017
Glass frog (Rulyrana spiculata) ventral / underside view, Cosnipata Valley, Peru
Abandoned dead and dying Orange trees that no longer have water to irrigate them near Bakersfield, California, USA. October 2014
Leaf insect (Phyllium sp. ) camouflaged in rainforest, Mulu National Park, BorneoLeaf insect (Phyllium sp.) camouflaged in rainforest, Mulu National Park, Borneo
Welwitschia (Welwitschia mirabilis) female plant with cones at sunset, Namib Naukluft National Park, Swakopmund, Namibia
Skull of male Urial sheep (Ovis vignei) on snow covered slope. Himalayas near Ulley, Ladakh, India
Greater flamingo (Phoenicopterus ruber) flock in flight, Pont de Gau, Camargue, France, May
Pecks skipper butterfly (Polites peckius) feeding on flowers, Crossways Preserve Montgomery County, Philadelphia, Pennsylvania, USA, June
Dogbane beetle (Chrysochus auratus) on dogbane, Crossways Preserve, Philadelphia, Pennsylvania, USA, August
Eastern chipmunk (Tamias striatus) gathering white oak acorns in cheek pouch, Chestnut Hill, Philadelphia, Pennsylvania, USA, September
Black-billed shrike-tyrant (Agriornis montanus) with nest material; Antisanilla Reserve, Ecuador
Violet-fronted brilliant (Heliodoxa leadbeateri) Ecuador
Green-crowned brilliant (Heliodoxa jacula) Mashpi Cloud Forest Reserve, Ecuador
Spiky flower-mimic stick insect (Toxodera berieri) active at night. Danum Valley, Sabah, Borneo
Urial sheep (Ovis vignei) herd running across steep barren slopes. Himalayas near Ulley, Ladakh, India
Lioness (Panthera leo) chasing away Cheetah (Acinonyx jubatus) Serengeti / Ngorongoro Conservation Area (NCA) near Ndutu, Tanzania
RF - Snow leopard (Panthera uncia) male moving over rocky, snow-covered slopes. Ulley Valley, Himalayas, Ladakh, India
Wolf (Canis lupus) male on rocky snow covered slopes. Ulley Valley in the Himalayas, Ladakh, India
Red-and-green macaw (Ara chloropterus) in flight over forest canopy. Buraco das Araras, Jardim, Mato Grosso do Sul, Brazil. September
Beach at the north of Principe Island, Democratic Republic of Sao Tome and Principe, Gulf of Guinea
White forest crab (Cardisoma armatum) portrait, seen through claw, Island of Principe UNESCO Biosphere Reserve, Democratic Republic of Sao Tome and Principe, Gulf of Guinea
Guinea lidless skink (Panaspis africanus) Principe Island, UNESCO Biosphere Reserve, Democratic Republic of Sao Tome and Principe, Gulf of Guinea
Blue shark (Prionace glauca) female off the coast of Portugal. August
Two Buff-tailed coronet (Boissonneaua flavescens) hummingbirds interacting, Andean montane forest, Ecuador. April
Two Cheetah cubs (Acinonyx jubatus) aged 12-14 days, Ngorongoro Conservation Area, Serengeti, Tanzania
Two Jaguars (Panthera onca) snarling at each other on the riverbank, Pantanal, Brazil. September
Herd of African Elephants (Loxodonta africana) grazing with cattle, Chobe National Park in Botswana
Sargassum fish (Histrio histrio) swimming with discarded fishing net, Dominica