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Nazca booby (Sula granti), pair billing on coastal cliff in evening light. Wolf Island, Galapagos. August 2016
Flightless cormorant (Phalacrocorax harrisi), adult and two chicks. Beagle Crater, Isabela Island, Galapagos
Blue-footed booby (Sula nebouxii), group fishing underwater. Itabaca Channel, Santa Cruz Island, Galapagos
Jaguars (Panthera onca) courting pair on a sand bank. Cuiaba River, Northern Pantanal, Mato Grosso, Brazil
Piraputanga, (Brycon hilarii) reflected on the water surface, Aquario Natural, Bonito, Mato Grosso do Sul, Brazil
Jaguar (Panthera onca) Pantanal, Mato Grosso, Brazil
Grass fire at night in Pantanal, Brazil
Kaieteur Falls is the worlds widest single drop waterfall, located on the Potaro river in the Kaieteur National Park, in Essequibo, Guyana, South America
Mount Roraima tepui (flat top mountain) is the highest of the Pakaraima chain of Tepuis plateaus in South America, Pakarajma Mountains, near Phillipai, West Guyana, South America
Galapagos racer snakes (Pseudalsophis biserialis) group alert watching for prey on beach, Galapagos, June. Endangered species. Taken on location for BBC tv Blue Planet 2 series
Galapagos giant tortoise (Geochelone nigra porteri), Cerro El Chato, Santa Cruz Island, Galapagos Islands
Brown booby (Sula leucogaster), adult diving for fish, Los Roques, Venezuela
Brazilian tapir (Tapirus terrestris) portrait, Tambopata Research Centre, Peru
Guianan cock-of-the-rock (Rupicola rupicola), two males fighting during lek, Guianan Shield, Suriname
Sol de Manana geyser bassin, Reserva Eduardo Avaroa, Altiplano, Bolivia
Puna / James flamingo (Phoenicoparrus jamesi) flock on Laguna Colorado, Reserva Eduardo Avaroa, Altiplano, Bolivia
Puma (Puma concolor) in high altitude habitat of Torres del Paine National Park, Chile
Llama standing in mud at the edge of Laguna Colorada, Bolivia
Giant cushion plant (Azorella compacta). Bolivia
High Altiplano with tussock grass called Paja brava (Festuca orthophylla) showing clonal growth spread. Bolivia. December 2016
Patagonian redfish (Sebastes oculatus) about 4cm, Comau Fjord, Patagonia, Chile
Julia heleconia (Dryas iulia) butterfly on head of Yacare caiman (Caiman yacare). Butterflies often land on caimans head to drink the salt from its eyes. Pantanal, Brazil
Spectacled caiman (Caiman crocodilus) in water at night, Mato Grosso, Pantanal, Brazil
Laguna 69, Huascaran National Park, Andes, Peru, November 2013
San Cristobal giant tortoises (Chelonoidis chatamensis), fighting, Galapaguera, San Cristobal Island, Galapagos
Marine iguana (Amblyrhynchus cristatus) in the surf, Black Beach, Floreana Island, Galapagos
Marine iguana (Amblyrhynchus cristatus) in water, Puerto Egas, Santiago Island, Galapagos
Galapagos lava lizard (Microlophus albemarlensis) sitting on Marine iguana (Amblyrhynchus cristatus) Punta Espinosa, Fernandina Island, Galapgos
Floreana giant tortoise hybrid descendants (Chelonoidis elephantopus) feeding on flower. These are descendants of the extinct Floreana giant tortoise, but are hybridised with other species
Galapagos green turtle (Chelonia mydas agassizii) swims over a coral reef. Darwin Island, Galapagos National Park, Galapagos Islands. East Pacific Ocean
Oceanic manta (Mobula birostris). Cousins Rock, San Salvador Island, Santiago Island, Galapagos National Park, Galapagos Islands. East Pacific Ocean
Galapagos barnacle blenny (Acanthemblemaria castroi) looking out from its home in an old barnacle shell. Cape Marshall, Isabela Island, Galapagos National Park, Galapagos Islands. East Pacific Ocean
Oreocallis grandiflora (Proteaceae family), Manu Cloud forest at 3500 metres altitude, Peru. November
Green-banded urania moth (Urania leilus) moths drinking salts from mineral-rich river clay, a behaviour know as puddling. On the banks of the Manu River, Manu Biosphere Reserve, Amazonia, Peru
Giant waxy monkey frog (Phyllomedusa bicolor). Lowland Amazon rainforest, Manu Biosphere Reserve, Peru
Three-striped poison frog (Ameerega trivittata) amongst leaf litter on lowland rainforest floor. Manu Biosphere Reserve, Amazonia, Peru. November
Nictitating membrane covering eye of Map Treefrog (Hypsiboas geographicus). The membrane affords camouflage and protection from dryness whilst allowing the frog to see
Andean Cock-of-the-rock male {Rupicola peruvianus} at lek in cloud forest canopy. Manu Biosphere Reserve, Amazonia, Peru. November
Brown / Tufted Capuchin (Cebus apella) family group in cloud forest, Manu Biosphere Reserve, Peru
Crab Spider (Thomisidae) waiting in an orchid to ambush its prey. Cloud forest, Manu Biosphere Reserve, Peru. November
Bush cricket or katydid (Tettigoniidae) with spines, camouflaged amongst moss. Forest interior of mid-altitude montane rainforest, Manu Biosphere Reserve, Peru
Hoatzin (Opisthocomus hoazin), pair on floating log in Cocha Salvador ox-bow lake. Manu Biosphere Reserve, Peru
RF - Red-and-green macaw (Ara chloropterus) flock feeding and flying. Wall of clay lick, Heath River, Tambopata / Bahuaja-Sonene Reserves, Amazonia, Peru / Bolivia border
Aerial view of lowland Amazonia rainforest in morning mist. Manu Biosphere Reserve, Amazonia, Peru. November 2017
Jaguar (Panthera onca palustris), male hunting amongst tree roots at edge of the Cuiaba River. Three Brothers, Porto Jofre, northern Pantanal, Mato Grosso, Brazil. August 2017
Marsh deer (Blastocerus dichotomus), stag. Pousada Araras, northern Pantanal, Mato Grosso, Brazil
Aerial view of Amazon River, with settlements and secondary rainforest, near Iquitos, Peru. July 2015
Scarlet macaw (Ara macao) in rainforest Tambopata National Reserve, Peru