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Whale shark (Rhincodon typus) close to the camera with a golden trevally fish (Gnathanodon speciosus) and outline of another whale shark behind, in the plankton-rich green waters of La Paz bay
Scarlet Macaws (Ara macao) drinking water inside an Amazonian claylick. Yasuni National Park, Orellana, Ecuador
Jaguar (Panthera onca) standing in mud, Yasuni National Park, Orellana, Ecuador
Marine Iguanas (Amblyrhynchus cristatus) basking. Fernandina Island, Galapagos Islands
Labiated rainfrog (Pristimantis labiosus) Mashpi, Pichincha, Ecuador
Black howler (Alouatta caraya) male and female, captive, occurs in Brazil and Paraguay. With digitally added leaf pattern
Red-footed booby (Sula sula) perched in tree. Genovesa Island, Galapagos
Nazca booby (Sula granti), pair on rocky shore at dawn. Punta Suarez, Espanola Island, Galapagos. May 2015
Nazca booby (Sula granti) standing on rock at coast. Punta Suarez, Espanola Island, Galapagos. October 2015
Blue-footed booby (Sula nebouxii) colony at sunet. Cape Douglas, Fernandina Island, Galapagos
Marine iguana (Amblyrhynchus cristatus), Punta Vicente Roca, Isabela Island, Galapagos
American flamingo (Phoenicopterus ruber) group of four with one displaying Punta Moreno, Isabela Island, Galapagos
Swallow-tailed gull (Creagrus furcatus) in flight, Plazas Islands, Galapagos
Flightless cormorant (Phalacrocorax harrisi) underwater, Punta Vicente Roca, Isabela Island, Galapagos, Ecuador, April
White-vented storm petrel (Oceanites gracilis galapagoensis) walking on surface of water whilst foraging, Galapagos
Marine iguana (Amblyrhynchus cristatus) on coast, Galapagos
Galapagos sea lions (Zalophus wollebaeki) fighting, Galapagos
Galapagos land iguana (Conolophus subcristatus) portrait with mouth open, Galapagos
Nazca booby (Sula granti) portrait, Galapagos, Ecuador
Great frigatebird (Fregata minor) male displaying inflated gular sac, Galapagos, Ecuador
Galapagos hawk (Buteo galapagoensis) landing, Galapagos, Ecuador. Vulnerable species
Galapagos hawk (Buteo galapagoensis) in flight, Galapagos, Ecuador. Vulnerable species
Galapagos hawk (Buteo galapagoensis) perched over sulphurous slope, Galapagos, Ecuador. Vulnerable species
People watching the Fissure eruption of Chico volcano into 9km diameter caldera. Sierra Negra volcano, Isabela Island, Galapagos, Ecuador. October 2005
Golden cownose rays (Rhinoptera steindachnyri) schooling in sheltered mangrove lagoon, Academy Bay, Santa Cruz Island, Galapagos, Ecuador
Salt cones on the Salar de Uyuni, Bolivia. March. The Salar is the worlds largest salt flat, at over 10500 square kilometres. Salt is shoveled into these cones, to be collected later. March 2012
James flamingo / Puna flamingo (Phoenicoparrus jamesi). Lago Colorado, Bolivia. March
Yacare caiman (Caiman yacare) Cuiaba River, Pantanal, Brazil, August
Yellow warbler (Setophaga petechia aureola) on lava rocks. Isabela Island, Galapagos. May
Great frigatebird (Phaethon aethereus) male at nest site, with gular sac inflated to attract females. Genovesa Island, Galapagos, May
Llama (Lama glama) Laguna colorada. Altiplano, Bolivia
Bolivian red howler (Alouatta sara), Pampas del Yacuma Protected Area, Bolivia
Hyacinth Macaw (Anadorhynchus hyacinthinus) two playing whilst hanging upside down, Pantanal, Brazil
Andean Emerald Hummingbird (Amazilia franciae), Mindo cloud forest area, Ecuador, July
Sword-billed hummingbird (Ensifera ensifera) in flight, North-Ecuador, Ecuador
Cotton-top tamarin (Saguinus oedipus), adult with two week old baby on back. Northern Colombia
Colombian red howler monkey (Alouatta seniculus) mother and baby in tree. Northern Colombia
Guanaco (Lama guanicoe) grazing on grassland plain, Patagonia, Chile. Novmeber
Llama (Lama glama) herd, Laguna colorada. Altiplano, Bolivia
Vicuna (Vicugna vicugna), five running across salt flat with mountain in background. Salar de Ascotan, Chile. September 2018
Andean fox (Lycalopex culpaeus) walking in the Altiplano, Andes, Bolivia. September 2018
Galapagos green turtle (Chelonia mydas agassizi) swimming in coastal waters of San Cristobal Island, Galapagos
Sergeant major (Abudefduf sp) schooling in mangroves of Isabela Island, Galapagos
Whistling heron (Syrigma sibilatrix sibilatrix), southern subspecies, native to Bolivia. Captive
Nazca booby (Sula granti), pair billing on coastal cliff in evening light. Wolf Island, Galapagos. August 2016
Flightless cormorant (Phalacrocorax harrisi), adult and two chicks. Beagle Crater, Isabela Island, Galapagos
Blue-footed booby (Sula nebouxii), group fishing underwater. Itabaca Channel, Santa Cruz Island, Galapagos
Jaguars (Panthera onca) courting pair on a sand bank. Cuiaba River, Northern Pantanal, Mato Grosso, Brazil