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Herd of wild Onaqui horses (Equus ferus) trotting in dust. Onaqui Mountain Herd Management Area, Utah, USA. September
Brown bear (Ursus arctos) walking across rock in forest, Land of the Leopard National Park, Russian Far East. Taken with remote camera. September
Sika deer (Cervus nippon) stag walking on mountain slope at the edge of woodland, with forested mountains in the background, Land of the Leopard National Park, Russian Far East
Sika deer (Cervus nippon) herd walking past fallen tree in forest as one looks around, Land of the Leopard National Park, Russian Far East. Taken with remote camera. June
Amur leopard (Panthera pardus orientalis) walking past snow-covered trees on mountain slope, Land of the Leopard National Park, Russian Far East. Critically endangered. Taken with remote camera
Sika deer (Cervus nippon) doe walking across snow covered mountain slope, Land of the Leopard National Park, Russian Far East. Taken with remote camera. December
Chocolate Labrador retriever puppy running on garden lawn, Rhode Island, USA. May
Humpback whale (Megaptera novaeangliae) creating bubble arc with fin after surfacing, Tubuai, French Polynesia, Pacific Ocean
Humpback whale (Megaptera novaeangliae) ascending to surface, Tubuai, French Polynesia, Pacific Ocean
Puma (Puma concolor) female, walking in deep snow, Torres del Paine National Park / Estancia Laguna Armarga, Patagonia, Chile
Puma (Puma concolor) cub, aged nine months, chasing scavenging White-throated Caracaras (Phalcoboenus albogularis), Torres del Paine National Park / Estancia Laguna Armarga, Patagonia, Chile
Three Guanacos (Lama guanicoe) running over hillside in front of the Towers granite rock formation and low cloud, Torres del Paine National Park, Patagonia, Chile
Puma (Puma concolor) female, running sequence in deep snow, Torres del Paine National Park / Estancia Laguna Armarga, Patagonia, Chile. Composite of five separate images
Two Sockeye / Red Salmon (Oncorhynchus nerka), female digging riverbed to lay eggs on spawning ground. Trees showing autumnal colours, Adams river, British Columbia, Canada. October
Two Leatherback turtle (Dermochelys coriacea) hatchlings moving across beach towards sea after leaving nest, Grande Riviere, Trinidad Island, Trinidad & Tobago, Caribbean Sea
Hanuman langur (Presbytis entellus) juvenile, running up tree trunk, Bandhavgarh National Park, India
Cockapoo puppy, aged 18 weeks, running, portrait
Cockapoo puppy, aged 4 months, running, portrait
Norfolk terrier puppy, aged 7 months, running, portrait
Galapagos sea lion (Zalophus wollebaeki) juvenile, jumping and playing in ocean, San Cristobal Island, Galapagos National Park, Pacific Ocean. Endangered
Puma (Puma concolor) cub running, Torres del Paine National Park, Magallanes, Chile
Herd of African elephants (Loxodonta africana) coming to drink at waterhole. Tsavo East National Park, Kenya
Wolf (Canis lupus) walking in forest, Finland. September
Amur / Siberian Tiger (Panthera tigris altaica) female in the wild, on a hillside, Lazovskiy zapovednik / Lazo Reserve protected area, Primorskiy krai, Far Eastern Russia, February 2012
Spinner dolphin (Stenella longirostris) juvenile surfacing next to its mother, Sri Lanka
Indri (Indri indri) male, leaping through the rain forest canopy, Andasibe-Mantadia National Park, eastern Madagascar. Critically Endangered
Wood duck (Aix sponsa) female, leading brood of ducklings on pond, Acadia National Park, Maine, USA. June
Black grouse (Lyrurus tetrix) male, jumping, part of a dominance display, at lek in early morning light, Kuusamo, Finland, April
Arctic fox (Vulpes lagopus) hunting along rocky shoreline under overhanging ice, Svalbard, Norway. July
Brown hare, (Lepus europaeus), male and female displaying courting behaviour, Islay, Scotland, UK. March
Brown hare, (Lepus europaeus), males in pursuit of female that is in season, Islay, Scotland, UK. March
Crested caracara (Caracara cheriway), adult, walking on beach. Pacific coast, Oaxaca state, Mexico. November
Group of Clarion angelfish (Holacanthus clarionensis) cleaning an Oceanic manta ray (Mobula birostris) as it swims over the reef, Revillagigedo Islands, Mexico, Pacific Ocean
Greater one-horned rhinoceros, (Rhinoceros unicornis) walking through shallow water, Bardia National Park, Terai, Nepal
Spotted deer (Axis axis), small herd leaping through grass, Bardia National Park, Terai, Nepal
Southern cassowary (Casuarius casuarius) walking on beach, Etty Bay, Queensland, Australia
Dung beetle (Scarabaeidae) rolling a ball of dung, Texas, USA. May
Common dolphin pod (Delphinus delphis) jporposing, Horta island, Azores, Atlantic Ocean
Alcedo giant tortoise (Chelonoidis vandenburghi) walking around caldera rim with fog rolling in during dry season, Alcedo Volcano, Isabela Island, Galapagos Islands, Ecuador
King penguins (Aptenodytes patagonicus) walk back to their breeding colony. Right Whale Bay, South Georgia Island
Red fox (Vulpes vulpes) vixen walking through snow, Derbyshire, UK. January
Mandarin duck (Aix galericulata) drake walking on snow-covered, frozen lake, London, UK. February
Black-capped chickadee (Poecile atricapillus) hopping between snow-covered perches, Lexington, Massachusetts, USA. February
Adelie penguin (Pygoscelis adeliae) leaping out of ocean as it approaches land, Devil Island, Antarctica, Southern Ocean
Red Lechwe (Kobus leche) herd crossing swamp surrounding a small island, Okavango Delta, Botswana, Africa
Family of Asian small-clawed otter (Aonyx cinerea), parent and pups, walking through grass. Captive, occurs in Asia. Zooparc Overloon, the Netherlands
Kerry bog pony, stallion, a rare breed, running, County Kerry, Republic of Ireland. April
Falabella miniature horse, stallion, walking over grassland, County Kerry, Republic of Ireland. April
