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Mute swan (Cygnus olor) parent and cygnets swimming. London, UK. May
Three juvenile Golden jackal (Canis aureus) pups running across a path, Danube Delta, Romania, July
Crab-plover (Dromas ardeola) three walking in a row on mudflat, two with food in beaks. Oman, June
Temmincks tragopan pheasant (Tragopan temminckii) Chongqing region, Sichuan, China
Arctic fox (Alopex lagopus) following scent, in winter pelage. Svalbard, Norway. April
Polar bear (Ursus maritimus), two cubs playing, leaping across sea ice, reflected in water. Svalbard, Norway, July
Polar bear (Ursus maritimus) walking across ice, snow covered mountain in background. Svalbard, Norway, April 2018
RF - Polar bear (Ursus maritimus) female walking across ice. Svalbard, Norway, April. (This image may be licensed either as rights managed or royalty free.)
Galapagos sea lion (Zalophus wollebaeki) jumping out of Pacific ocean. Sombrero Chino, Santiago Island, Galapagos
RF - Puma (Puma concolor puma), young male walking on hillside near Torres del Paine National Park, Patagonia, Chile. November. (This image may be licensed either as rights managed or royalty free.)
Grizzly bear (Ursus arctos) running through water, Lake Clark National Park, Alaska, September
Eastern white-bearded wildebeest (Connochaetes taurinus), four running in line, on migration, Masai Mara National Reserve, Kenya
Bengal tiger (Panthera tigris) running down slope. Ranthambore National Park, India. Photo© Phillip Ross/Felis Images
American alligator (Alligator mississippiensis) emerging from water, in evening light. Myakka River State Park, Florida, USA. March
Gentoo penguins (Pygoscelis papua) walking in line, returning to nesting area, Port Charcot, Antarctic Peninsula, Antarctica
Red squirrel (Sciurus vulgaris) foraging in forest, climbing down tree trunk, Cairngorms National Park, Scotland, October
RF - Brown Hare (Lepus europaeus) running through field of grass, Scotland, UK.May (This image may be licensed either as rights managed or royalty free.)
Vicuna (Vicugna vicugna), five running across salt flat with mountain in background. Salar de Ascotan, Chile. September 2018
Andean fox (Lycalopex culpaeus) walking in the Altiplano, Andes, Bolivia. September 2018
Black Labrador dog, age 6 months, walking
Red squirrel (Sciurus vulgaris) jumping towards camera. Scotland, UK. February
Wild Grey wolf (Canis lupus) walking, Saxony-Anhalt, Germany
Jaguar (Panthera onca) Pantanal, Mato Grosso, Brazil
Double-crested basilisk (Basiliscus plumifrons) running across water surface, Santa Rita, Costa Rica
Gentoo penguin (Pygoscelis papua) walking back to the colony, Sea Lion Island, Falkland Islands, October
White-headed capuchin (Cebus capucinus) walking on giant palm-tree, Costa Rica
Thorny dragon (Moloch horridus) in desert habitat, Australia
Namib sand snake (Psammophis namibensis) in sand dunes, Swakopmund, Erongo Region, Namibia
Jaguar (Panthera onca) male and female four month old cubs playing, native to Southern and Central America, captive
Fluffy black kitten, 10 weeks, walking
Leopard (Panthera pardus) climbing in tree. Kabini, Nagarhole National Park, Karnataka, India
Shetland sheepdog running in Common bluet (Houstonia caerulea) field. Haddam, Connecticut, USA. May
Lion (Panthera leo), female jumping over a stream. Mana Pools National Park, Zimbabwe
Snowy owl (Bubo scandiacus) walking on ground in snow, one foot raised, Ontario, Canada, January
Bottlenosed dolphin (Tursiops truncatus) porpoising during annual sardine run, Port St Johns, South Africa. June
RF - Lion (Panthera leo) male running in desert, Kgalagadi Transfrontier Park, South Africa
RF - Siberian tiger (Panthera tigris altaica) in snow, captive
Black Merens stallion running in pasture. Northern France, Europe. February
Bay Icelandic horse trotting in the snow, Snaefellsnes Peninsula, Iceland, March
Golden pheasant (Chrysolophus pictus) male jumping, Yangxian Biosphere Reserve, Shaanxi, China
White eared pheasant (Crossoptilon crossoptilon) spreading his wings while jumping down a rock, Baima Snow Mountain Nature reserve, Yunnan, China, October
Atlantic salmon (Salmo salar) jumping up waterfall during spawning migration upstream, Ume River, Sweden. July. Photographed for The Freshwater Project
Kerry Hill domestic sheep, spring lambs running, England, UK
Grey wolf (Canis lupus) with Saiga antelope (Saiga tatarica) in the background, Astrakhan Steppe, Southern Russia
Polar bear (Ursus maritimus), adult male, outside Kaktovik, Alaska, USA
Raccoon dog (Nycterentes procyonoides) walking across snow, Vladivostok, Primorsky Krai, Far East Russia. November
Eurasian lynx (Lynx lynx) walking in snow, Yaroslavl, Central Federal District, Russia. March
Smooth-coated otter (Lutrogalle perspicillate) about to enter a marsh at dawn, Dudhwa National Park, Uttar Pradesh, India. Photographed using a camera trap