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RF - Coquerels sifaka (Propithecus coquereli) jumping, Anjajavy Private Reserve, north west Madagascar
Great hammerhead shark (Sphyrna mokarran) in shallow water. South Bimini, Bahamas. The Bahamas National Shark Sanctuary. Gulf Stream, West Atlantic Ocean
Glorious scarab (Chrysina gloriosa) flying, Brewster County, Texas, USA. Controlled conditions. August
Common frog (Rana temporaria) portrait, Broxwater, Cornwall, UK. June
Small tortoiseshell butterfly (Aglais urticae) resting on old painted door. Dorset, UK, August
Black-headed gull (Chroicocephalus ridibundus) alighting on water at dawn, Cheshire, UK, November. Highly Commended in the British Wildlife Photography Awards (BWPA) competition 2013
RF- Small jellyfish (Pelagia noctiluca) swimming beneath surface. Marine Protected Area of Portofino (Area Marina Protetta, Portofino), Liguria, Italy. Mediterranean Sea
Burrowing owl (Athene cunicularia) fledgling with head on side, Pantanal, Brazil. November
Luminescent jellyfish / Mauve stinger (Pelagia noctiluca) in open water, Gozo Island, Malta. Mediterranean Sea
Queen Marys Tree, a veteran Sweet chestnut (Castanea sativa) over 600 years old, Cumbernauld, Scotland, UK, July
Great egret (Ardea alba) flying across lake, Ranthambhore National Park, India
Arctic tern (Sterna paradisaea) in flight with fish in beak for feeding chick, Machias Seal Island, Bay of Fundy, New Brunswick, Canada, July
Three Strawberry poison frogs (Oophaga pumilio) on rock, Sarapiqui, Heredia, Costa Rica
Sword billed hummingbird (Ensifera ensifera) in flight, Guango, Napo, Ecuador
Group of African elephants (Loxodonta africana) silhouetted at sunrise, Okavango Delta, Botswana
Red fox (Vulpes vulpes) rear view sitting on grass close to its den, Derbyshire, UK
Dusky shark (Carcharhinus obscurus) swimming through light rays during sardine run. Wild Coast, Eastern Cape, South Africa. June
Maned wolf (Chrysocyon brachyurus) setting out to hunt in grassland at dusk. Serra da Canastra National Park, Brazil
Sparrowhawk (Accipiter nisus) juvenile flying, Parainen Uto, Finland. September
Annas hummingbird (Calypte anna) male dancing in flight and displaying his bright pink throat feathers to a female, Montana, USA. June
Snakelocks anemone (Anemonia viridis) growing on a stalk of Eelgrass (Zostera marina) on the seabed, Isle of Coll, Inner Hebrides, Scotland, UK, Atlantic Ocean. August
Lesser Egyptian jerboa (Jaculus jaculus) in the desert at night, Oued Afra, Western Sahara, Africa
Female scalefin anthias (Pseudanthias squamipinnis) swimming in front of a school of glassfish (Apogonidae), whose shiny bodies are rendered into out of focus bokeh patterns. Laamu Atoll, Maldives
Mute swan (Cygnus olor) swimming on sun-dappled water, Lower Silesia, Poland
Mandarin duck (Aix galericulata) drake standing on frozen pond during snowstorm. Richmond Park, London, UK. January
Male Black woodpecker (Dryocopus martius) in snow, showing red crown. Kuusamo Finland December. Overall winner of the Swiss Vogelwarte photo competition 2021
Pygmy leaf-folding frog (Afrixalus brachycnemis) sitting on a fern frond, Tanzania, Africa
Song thrush (Turdus philomelos) looking for worms in a garden, Broxwater, Cornwall, UK. April
Bearded vulture (Gypaetus barbatus) flying, Ordesa National Park, Pyrenees, Spain
Thorny devil stick insect (Eurycantha calcarata), Willaumez Peninsula, New Britain, Papua New Guinea, December
Grey triggerfish (Balistes capriscus) portrait, Tenerife, Canary Islands. August
Portrait of a rescue dog on white background, one ear perked up
Buzzard (Buteo buteo) landing on branch, winter, Lorraine, France, February
Rosemary beetle (Chrysolina americana) feeding on Lavender (Lavandula angustifolia) in garden Cheshire, UK, June
RF - Sailing boat in mist at sunrise, reflected in Wimbleball Lake. Exmoor National Park, Somerset, England, UK. September 2020. (This image may be licensed either as rights managed or royalty free.)
RF - Small tortoiseshell butterfly (Aglais urticae) nectaring on Verbena (Verbena sp). Cornwall, England, UK. September. (This image may be licensed either as rights managed or royalty free.)
Scoophead shark (Sphyrna media), released specimen. Isla Chepillo, Rio Bayano, Panama
Cuban dogfish (Squalus cubensis). Exuma Sound, Bahamas
Atlantic sixgill shark (Hexanchus vitulus) swimming with open mouth. Cape Eleuthera, Bahamas
Blotched blue-tongue skink (Tiliqua nigrolutea) female gaping with tongue out, in defense against perceived threat. Lake Tantangara, Snowy Mountains, New South Wales, Australia
Mute swan (Cygnus olor) stretching its wings after preening on a misty morning. Valkenhorst Nature reserve, Valkenswaard, The Netherlands, May
Arctic fox (Vulpes lagopus), blue colour morph in winter coat walking through snow. Hornstrandir Nature Reserve, Iceland. February
Peregrine falcon (Falco peregrinus) in flight with open beak. The Netherlands. May
Red-and-green macaw (Ara chloropterus) in flight, Pantanal, Brazil. August
Common tern (Sterna hirundo) flying low over water. Hjalstaviken, Uppland, Sweden
Arabian gazelle (Gazella gazella), Jaluni, Oman, May
Kottigehar dancing frog (Micrixalus kottigeharensis), male waving foot and calling, territorial behaviour. Agumbe, Western Ghats, India. Endemic
European red fox (Vulpes vulpes) peeking out of a snow bank. Gran Paradiso National Park, Italy