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Mute swan (Cygnus olor) on a misty morning, with wings raised up in aggressive stance, Valkenhorst Nature Reserve, Valkenswaard, the Netherlands. December
Indian Flying Fox (Pteropus giganteus), flying, India
Greater roadrunner (Geococcyx californianus), adult drinking, Rio Grande Valley, South Texas, Texas, USA. June
Snowy owl (Bubo scandiacus) on beach. Ocean County, Washington, USA. March
Ruddy turnstone (Arenaria interpres) in breeding plumage. Yukon Delta National Wildlife Refuge. Alaska, USA. June
Juvenile male Peregrine falcon (Falco peregrinus) in flight chasing his parent who has Feral pigeon (Columba livia) kill in claws, Bristol, England, UK, June
Silhouetted Dartmoor pony (Equus caballus) at sunrise, Combestone Tor, Dartmoor National Park, Devon, England, UK, January
RF- Small group of Pink-footed geese (Anser brachyrhynchus) in flight at dawn, The Wash Estuary, Norfolk, England, UK, October 2011
RF- Chaffinch (Fringilla coelebs) male in flight in snow. Glenfeshie, Scotland, February
RF- Rear view of Basking shark (Cetorhinus maximus) feeding on plankton, visible as white dots on water surface near island of Coll, Inner Hebrides, Scotland, UK. June
Red deer (Cervus elaphus) stag in heavy snow, Scotland, UK, February
Black grouse (Tetrao tetrix) male in flight, Cairngorms National Park, Scotland, UK, April
Grey seal (Halichoerus grypus) swimming over kelp forest, Shetland, Scotland, UK, August
Black-throated diver (Gavia arctica) on water, Finland, May
Goldfinch (Carduelis carduelis) adult feeding on teasel seeds, UK, February
European Greenfinch (Chloris chloris) flying at dawn. When backlit the tiny transparent barbs that make up the feather act like a prism revealing vivid colours in the early light
European honey bee (Apis mellifera) in flight, feeding on flowers (Brachyglottis sp). Monmouthshire, Wales, UK, June
Black veined white butterfly (Aporia crataegi) on grass, Gavarnie-Gedre, Pyrenees National Park, France, June
Rough seas at Creac h lighthouse during Storm Ruth, Ile d Ouessant, Armorique Regional Park, Iles du Ponant, Finistere, Brittany, France, Iroise Sea. 8th February 2014
Tree sparrows (Passer montanus) perching on a branch in the rain. Perthshire, Scotland, December
White-tailed eagle (Haliaeetus albicilla) in flight, Norway, April
Bald eagle (Haliaeetus leucocephalus) head portrait, Alaska, USA, February
Blackbird (Turdus merula) male in flight during snowfall, Oisterwijk, The Netherlands. January
Red deer (Cervus elaphus) stag close up in the snow, Scotland, March
Large group of Cavalier King Charles Spaniel, puppies, three of which have tricolour coat and the rest blenheim
Blue-spotted dwarf iguana (Enyalioides praestabilis) on branch, Sumaco, Napo, Ecuador
Las gralarias glass frog (Nymphargus lasgralarias) on leaf, Mindo, Pichincha, Ecuador
Amazon wood lizard / Guichenots dwarf iguana (Enyalioides laticeps) portrait, Yasuni National Park, Orellana, Ecuador
Shieldhead gecko (Gonatodes caudiscutatus) on branch, Canande, Esmeraldas, Ecuador
Eyelash viper (Bothriechis schlegelii) on twig, Canande, Esmeraldas, Ecuador
Pinocchio lizard (Anolis proboscis) male on twig, Mindo, Pichincha, Ecuador, January 2013, Endangered species
Southern yellow billed hornbill (Tockus leucomelas) perched on branch, Etosha National Park, Harare Province, Namibia
Green jararaca / Two striped forest pit viper (Bothriopsis bilineata) on branch, Tambopata, Madre de Dios, Peru
Tiger leg monkey frog / Tiger striped monkey frog (Phyllomedusa tomopterna) portrait, on stem, Yasuni National Park, Orellana, Ecuador
Snowy owl (Bubo scandiacus) in flight, Boundary Bay, British Columbia, Canada. February. First place in the Overall Category of the 2013 Great Backyard Bird Count photo competition
Steppe eagle (Aquila nipalensis) in flight, Oman, November
RF- White tailed sea eagle (Haliaeetus albicilla) in flight, Flatanger, Norway, June
Ash Tree (Fraxinus excelsior) growing in a field. Peak District National Park, Derbyshire, UK. August
Mountain hare (Lepus timidus) grooming itself, with back foot raised, Cairngorms National Park, Scotland, UK, February. Highly Commended in the Animal Behaviour category of the British Wildlife
Red deer (Cervus elaphus) female doe portrait, Jura, Scotland, UK, September
Water vole (Arvicola amphibius) feeding, Kent, UK May
RF - Female emperor dragonfly (Anax imperator) laying eggs on garden pond, Broxwater, Cornwall, UK. July
RF- Common frog (Rana temporaria) portrait with red background. UK
RF- Little Owl (Athene noctua) perched on tree stump covered in moss. Worcestershire, England, UK. May 2013
Male Yellow-headed jawfish (Opistognathus aurifrons) blows out the clutch of eggs he is incubating in his mouth to oxygenate them. East End, Grand Cayman, Cayman Islands, British West Indies
Gemsbok (Oryx gazella) aerial view of herd on sand, Namib Desert, Namibia
Arabian Oryx (Oryx leucoryx) at sunset. Previously extinct in the wild, their conservation status is now stable thanks to conservation efforts.This male is part of a reintroduction program in Dubai
Toco Toucan (Ramphastos toco) head and beak profile portrait, Brazil
