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Common toad (Bufo bufo) on Brandon Hill with Cabot Tower in the background, Bristol, UK. March
White-throated Mountain Gem (Lampornis castaneoventris), Green-crowned Brilliant (Heliodoxa jacula) group of three with one in flight, Savegre, Costa Rica
Leaf cutter ants (Atta sp) carrying plant matter, Costa Rica
Solar eclipse, Finland, 20 March 2015
Downy emerald dragonfly (Cordulia aenea) in flight, Joutsa, Finland, July
Silver washed fritillary butterfly (Argynnis paphia) female in flight with thistles, Uusimaa, Finland, August
Black-and-white kitten, Solo, 6 weeks, taking a selfie. Composite image
Northern gannets (Morus bassanus). Bass Rock, Scotland, UK. August
View up to Beech (Fagus sylvatica) woodland canopy in winter, photographed with a circular fisheye lens. Derbyshire, UK. December
Pebbles reflected in the water surface at mouth of Sheryldy river, Lake Baikal, Siberia, Russia. September
European brown bear (Ursus arctos) head portrait, captive, occurs in Eurasia
Mute swan (Cygnus olor) flock on snow, Hazerswoude, The Netherlands, February
Golden eagle (Aquila chrysaetos) portrait, captive, occurs in the Northern hemisphere
Robin (Erithacus rubecula) profile in field studio, Scotland, UK, January
Black and white photograph of saltmarsh at Annat, Beinn Torridon, Scotland, UK, November
Grey wagtail (Motacilla cinerea) at waters edge, Gaoligong Mountain National Nature Reserve, Yunnan Province, October
Whooper swans (Cygnus cygnus) in misty hot springs, Lake Kussharo, Hokkaido, Japan, February
Tsaranoro massif, Andringitra NP, Madagascar, October 2015
Red-backed shrike (Lanius collurio) adult male taking off, Lower Saxony, Germany, June. Sequence 2 of 3
Atlantic Puffin (Fratercula arctica) covered by mud from its partner digging out their nesting chamber / burrow, Skomer Island, Wales, UK, April
Young California sea lion (Zalophus californianus) play in the canopy of a forest of Giant kelp (Macrocystis pyrifera). Santa Barbara Island, Channel Islands. California, USA. North East Pacific Ocean
Wolf fish (Anarhichas lupus) portrait, Akuyeri, Iceland. North Atlantic Ocean
Lion (Panthera leo) male sharpening his claws on a tree at night, Sabi Sand Game Reserve, South Africa
Prayer plant (Calathea plowmanii) close up of leaf, TU Delft Botanical Garden, Netherlands, August
Aerial view of eroded banks and river with suspended load, caused by deforestation, central Madagascar
European brown bear (Ursus arctos) scratching up against tree in snow, Finland, April
Black bear (Ursus americanus) male close up head portrait, Minnesota, USA, June
Coyote (Canis latrans) jumping in snow with its prey, Yellowstone National Park, Wyoming, USA, February
Head portrait of grey Andalusian stallion in dark barn, Northern France, Europe
Grey Arabian stallion running in desert dunes near Dubai, United Arab Emirates
Two wild Mustang stallions fighting in White Mountain Herd Area, Wyoming, USA. August
Wild Mustang stallion chasing another away from his family in White Mountain Herd Area, Wyoming, USA. August
European beaver (Castor fiber) foraging at dusk, Knapdale Forest, Argyll, Scotland. Highly commended in the BWPA (British Wildlife Photography Awards) Competition 2016
Red Grouse (Lagopus lagopus scoticus) male on rock in swirling spindrift of snow, Scotland, UK, February. Highly commended in the Habitat category of the BWPA
Sick sponge, stricken by a Cyanobacteria, Lake Baikal, Siberia, Russia. December
Cheetah (Acinonyx jubatus), cubs age 8 weeks playing, Masai Mara Game Reserve, Kenya
Common toad (Bufo bufo) pair in amplexus, Deeside, Scotland April
Silky shark (Carcharhinus falciformis) San Benedicto Island, Revillagigedo Archipelago Biosphere Reserve, Socorro Islands, Western Mexico
Head portrait of grey Andalusian stallion running with mane flowing in arena in Northern France, Europe
Illustration of Hornets and Wasps, from Arcana entomologica, or, Illustrations of new, rare, and interesting insects by J.O. Westwood, 1845
Landscape of ice on Lake Baikal, Siberia, Russia, Lake Baikal, Siberia, Russia. March 2015
Auckland Island brown teal (Anas aucklandica) flightless and feeds and roosts in onshore kelp beds, lives predominantly on Enderby Island, Auckland Islands, Subantarctic New Zealand January
Southern hawker dragonfly (Aeshna cyanea) flying in habitat, Joutsa, Central Finland, August
Atlantic Puffin (Fratercula arctica) swimming, split level view, Skomer Island, Wales, UK, July
Atlantic Puffins (Fratercula arctica) in flight in mist, Skomer Island, Wales, UK, August
North Pacific right whale (Eubalaena japonica) swimming near surface, aerial view, Channel Islands National Park, California, USA. February
Avalanche in the mountains of Torres Del Paine National Park, Southern Patagonia, Chile
Orchid (Bulbophyllum sp) on white background, Botanic Garden Leiden, Netherlands, July
