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Red-winged blackbird (Agelaius phoeniceus) male in territorial display, Virginia, USA. May
Blackspotted stingray (Taeniurops meyeni) with a remora (Echeneis naucrates) attached to its belly swimming in the open water with a diver in the background. Ari Atoll, Maldives, Indian Ocean
Chocolate chip sea star o (Protoreaster nodosus) on sandy bottom. Mabul, Malaysia
Bacteria (Leptothryx discophora) causing iridescent sheen on water surface with White willow leaf (Salix alba). Sierra de Grazalema Natural Park, southern Spain, November
Poplar trees (Populus sp) in autumn, Sierra de Grazalema Natural Park, southern Spain, October
Hawthorn berries (Crataegus monogyna) in Poplar woodland (Populus) Sierra de Grazalema Natural Park, southern Spain. October
Frozen Elmleaf blackberry (Rubus ulmifolius) in a small pond, Sierra de Grazalema Natural Park, southern Spain, January
Fly agaric (Amanita muscaria) in foggy forest, Los Alcornocales Natural Park, southern Spain, January
Leaf frozen in ice, Sierra de Grazalema Natural Park, southern Spain, February. Highly Commended in the Plants and Fungi Category of the Asferico 2016 Awards
Head portrait of grey almost white Andalusian mare running in snow, Berthoud, Colorado, USA. January
Asiatic elephants (Elephas maximus), dust bathing at dawn. Jim Corbett National Park, India
Kottigehara dancing frog (Micrixalus kottigeharensis), dancing frog name given because of foot flagging behavior between competing males. Endemic to Western Ghats. Agumbe, Karnataka, India
Jayarami Bush frog (Raorchestes jayarami), inflating vocal sac, calling to attract mate, Endemic to Western Ghats. Anamalai Wildlife Sanctuary, India
Ocelot (Leopardus pardalis) portrait. camera trap image, Nicoya Peninsula, Costa Rica
Polar bear (Ursus maritimus) aerial view of sow with cub along the Arctic coast in summer, near Point Barrow, Alaska, USA. July
Black-crested titmouse (Baeolophus bicolor), adult perched on icy branch of Possum Haw Holly (Ilex decidua) with berries, Hill Country, Texas, USA. February
Tricolour merle Collie puppy, Indie, age 10 weeks
Sea stack and Talisker Beach, Isle of Skye, Inner Hebrides, Scotland, UK, September 2013
Scots pine (Pinus sylvestris) forest in evening light, Abernethy, Cairngorms National Park, Scotland, UK, June
Bison (Bison bison) close up head portrait with snow on fur, Yellowstone National Park, USA, February
Pine marten (Martes martes) leaping, Ardnamurchan Peninsula, Scotland, August. Highly commended in the Animal Behaviour category of the BWPA (British Wildlife Photography Awards) Competition 2016
Gannet (Sula bassana) in water, Shetland, UK. July
Kestrel (Falco tinnunculus) male, Hampstead Heath, England, UK
Red kite (Milvus milvus) in a dive, Gigrin Farm, Rhayader, Wales, UK April
Studio image of multiple Nudibranch species. Gulen, Norway. North East Atlantic Ocean. Digital composite
School of Blacksmith fish (Chromis punctipinnis) swim through a giant kelp (Macrocystis pyrifera) forest. Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America
Nassau grouper (Epinephelus striatus) yawning on a coral reef drop off. San Salvador island, Bahamas. Bahamas Sea, Tropical West Atlantic Ocean
Napoleon wrasse (Cheilinus undulatus) portrait. Red Sea, Egypt. October
Cirrus clouds displaying wind shear, Brechin, Scotland, UK, September
Monte Perdido massif and Las Tres Sorores (The Three Sisters) Ordesa National Park, Aragon, Huesca, Spain
Portrait of an African elephant (Loxodonta africana) on Selinda Reserve, Botswana
Common dolphin (Delphinus delphis) reflection as its swimming on the surface of the ocean, Eastern Pacific, March
Sperm whales (Physeter macrocephalus) large group gathering together, rubbing together to shed skin, interacting and defecating. Indian Ocean, March
Willow grouse (Lagopus lagopus) camouflaged against snow, Inari, Kiilopaa, Finland, February
Bay of Laig, Isle of Eigg towards Isle of Rum, Inner Hebrides, Scotland, UK, April 2014
Sedge warbler (Acrocephalus schoenobaenus) singing with bokeh effect in background, Norfolk, England, UK, May. Winner of the Birds in Environment Category of the Bird Photographer of the Year 2015
Natural ice sculpture floating at sea in Svalbard, Norway
Panoramic landscape of Cerredo mountain at sunset, Castro Urdiales, Cantabria, Spain
Common / Northern minke whale (Balaenoptera acutorostrata) surfacing, Pacific coast of Baja California, Mexico, February
Grayling (Thymallus) under Ice, Lake Baikal, Siberia, Russia
Royal spoonbill (Platalea regia) preening and shaking head feathers, Victoria, Australia
Grey heron (Ardea cinerea) in rain with another heron behind, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Two Buzzards (Buteo buteo) fighting, Lake Csaj, Kiskunsagi National Park, Hungary. February
Cormorants (Phalacrocorax carbo) squabbling over fish, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Two Burrowing owls (Athene cunicularia) fledgling with adult about to take off, Pantanal, Brazil
Grey herons (Ardea cinerea) fighting, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Grey heron (Ardea cinerea) squabbling, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. February
Shadowfin soldierfish (Myripristis adusta) West Papua, Indonesia