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Twelve Apostles rock formations, Great Ocean Road, Port Campbell National Park, Victoria State, Australia, September, 2011
Lower peaks of the Mont Cervin / Monte Cervino / The Matterhorn massif, Pennine Alps, Switzerland / France border, April 2011
Common / Edible cockle (Cerastoderma / Cardium edule) shells on beach, Belgium
Corcovado Hill and the Christ Redemptor statue in the Atlantic Rainforest of Tijuca National Park, Rio de Janeiro City, Rio de Janeiro State, Southeastern Brazil, August 2011
Harvest mouse (Micromys minutus soricinus) feeding on blackberry, captive, Dorset, UK, October
German Hound male, 6 months
Mourtia beach and bay with the southeast tip of Samos and Mount Mycale in Turkey sMourtia beach and bay with the southeast tip of Samos and Mount Mycale in Turkeys Dilek Peninsula National Park in the background, east coast of Samos, August 2011
Honeybee workers exchanging food - known as trophallaxis (Apis mellifera) Sussex, UK
Mount Pico, volcanic mountain, Pico Island, Azores, March, 2010
Sparkling violetear hummingbird (Colibri coruscans) at base of Chimborazo Volcano (Highest mountain in Ecuador), Andes, Ecuador, South America
Caribbean reef shark (Carcharhinus perezi) Jardines de la Reina National Park, Cuba
Jersey tiger moth (Euplagia quadripunctaria) large numbers flying to and resting on Sweetgum trees (Liquidambar orientalis) in the Petaloudes Valley, Rhodes island, Greece, August 2011
Row of Starlings (Sturnus vulgaris) perched on wire. Marais Breton, Vendee, French Atlantic Coast, November
Acacia sp. at night, with a herd of Plains Zebra (Equus quagga) beneath. Near Ndutu, Nogorongoro Conservation Area, Serengeti National Park, Tanzania
Red waters of the Rio Tinto, coloured by dissolved minerals, primarily iron. Andalusia, Spain, December 2011
Brown Long-eared bat (Plecotus auritus) drinking from a lily pond, Surrey, UK. Overall WINNER of the Audubon Society of Greater Denver Share the Viewae competition 2011
Common dolphins ( Delphinus delphis) swimming below the surface, Atlantic Ocean, Portugal, October
A pod of Bottlenose dolphins (Tursiops truncatus) swimming into the Sado river, Portugal, July
Crowned crane (Balearica regulorum) close-up of feathers, captive
Greater flamingo (Phoenicopterus ruber) close-up of back, captive
Male Indian peafowl (Pavo cristatus) displaying, captive
Black swan (Cygnus atratus) captive
Two Scarlet macaws (Ara macao) flying, captive
Ginger Kittens (Felis catus) in garden. France, September
Peyreleau medieval village. Gorge de la Jonte, Causse, Cevennes, Massif Central, Ayeyron, France, September 2011
Comtois Horse (Equus caballus). Camargue reserve, Rhone, France, September
Iceberg off the Antarctic Peninsula, Antarctica, February 2009, Taken on location for BBC Frozen Planet series
Male Whinchat (Saxicola rubetra) perched on Oil seed rape (Brassica napus) in a field, Vosges, France, April
Roe deer (Capreolus capreolus) at sunset, Vosges, France, August
Tropical Butterfly (Adepha) on a leaf with wings spread. Danum Valley, Sabah, Borneo, Malaysia
Agave (Agave sp. ) leaf close-up detail. Maui, Hawaii, FebruaryAgave (Agave sp.) leaf close-up detail. Maui, Hawaii, February
Ichneumon wasp (Compsocryptus sp. ?) in flight, Williamson county, Texas, USA, NovemberIchneumon wasp (Compsocryptus sp.?) in flight, Williamson county, Texas, USA, November
Composite image of flying insects visiting blacklight, Texas, USA, September
House sparrows (Passer domesticus) perched on building, Morocco, March
Limber Pine (Pinus flexilis) against high mountain peaks. Grand Teton National Park, Wyoming, USA. Autumn 2010
Limber Pine (Pinus flexilis). Grand Teton National Park, Wyoming, USA, October 2010. Famous patriarch tree of the Grand Tetons just after fall snowstorm
Bridled nailtail wallaby (Onychogalea fraenata) Queensland, Australia, Endangered species, October
Small copse of trees under a summer sky. Champagne, France, July
Lone Walnut (Juglans) tree in a field of barley. Picardy, France
Oak (Quercus) covered with frost. Picardy, France
Portrait of a Lynx (Lynx lynx) mother and kitten. Hanau, Germany, July. Captive
Dipper (Cinclus cinclus) perched on moss-covered waterfall, Peak District NP, Derbyshire, UK, May, Highly Commended, Habitat category, British Wildlife Phototgraphy Awards (BWPA) competition 2011
Snow goose with gosling (Chen caerulescens) at sunset. Taken on location for BBC Frozen Planet series, 2008
Southern elephant seal (Mirounga leonina) close up of sleeping individual, South Georgia. Taken on location for BBC Frozen Planet series, 2008
Marbled White Butterfly (Melanargia galathea) resting on grass stem, Devon, UK July. Winner of Seasons category of the BWPA 2011 competition
Great Blue Shark (Prionace glauca), dorsal view against dark water. Over 20 million Blue Sharks are fished annually, as bycatch and due to the high value of their fins
Brown hare (Lepus europaeus) running past hay barn in snow storm, Lapua, Finland, March. WINNER, Animals in their Environment category, 2011 WILDLIFE PHOTOGRAPHER OF THE YEAR competition
Pyramid of sea ice catching the first rays of the returning sun in the high Arctic, Svalbard, Spitsbergen, Norway, February 2009. Taken on location for the BBC series, Frozen Planet
