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Cotton Grass (Eriophorum vaginatum) growing in water. Goldenstater Moor, Niedersachsen, Germany
Great Tit (Parus major) in flight. Bavaria, Germany, December
Male Andean Cock-of-the-Rock (Rupicola peruvianus) displaying. Cock-of-the-Rock Lodge, Peru, August
Looking up through the leaves of Wild sarsaparilla (Aralia nudicaulis) in woodland, Cape Cod, Massachusetts, USA, May
Lava field with growth of Dotted polypody fern (Polypodium pellucidum), Kilauea, Hawaii, USA, August 2008
A wild newborn colt (Equus caballus) standing up being nuzzled by its mother. Foothills of Reno, Nevada, USA, May
Pine marten (Martes martes) in tree trunk, Finland, July
Two Common Bottlenose Dolphin (Tursiops truncatus) in shallow water. Punta Allen, Sian Ka an Biosphere Reserve, Quintana Roo, Yucatan Peninsula, Mexico
Thousand-year-old Canary Juniper, (Juniperus turbinata canariensis / Juniperus phoenicea var)Thousand-year-old Canary Juniper, (Juniperus turbinata canariensis/ Juniperus phoenicea var. turbinata ) against a dusky sky. El Sabinar, El Hierro Island, Canary Islands, February 2011
The Lovers Baobabs (Adansonia grandidieri). Near Morondava, Madagascar" The Lovers" Baobabs (Adansonia grandidieri). Near Morondava, Madagascar, Africa, September 2010
Isolated rain storm above the sea. Bahamas Sea, Bahamas, North America, August 2006
Rough coated Jack Russell Terrier puppy, black, tan and white, portrait
Rough coated Jack Russell Terrier, tan and white, lying down, panting
Aerial view of the rivers and saltmarshes of the Bahia / Bay de Cadiz Natural Park, Andalucia, Spain, March 2008
Dawn over the Segama River, with mist hanging over lowland rainforest. Heart of Danum Valley, Sabah, Borneo
Giraffe (Giraffa camelopardalis) female bending down to calf, Okavango Delta, Botswana
Unravelling Fern fronds (Unknown species). Near Nepenthes Field Camp, mid-altitude montane forest. Centre of Maliau Basin, Sabahs Lost World, Borneo
Asiatic lion cub (Panthera leo persica) looking up into tree, possibly at a bird, Gir Forest NP, Gujarat, India
Birch Sawfly (Cimbex femoratus) feeding on a leaf. August, UK
Dalmatian pelicans (Pelecanus crispus) low angle perspective of open bills, Lake Kerkini, Greece, February. Highly Commended, 2011 Wildlife Photographer of the Year competition
Cane Toad / Marine Toad / Giant Toad (Bufo marinus) adult jumping in Dogweed (Dyssodia pentachaeta) field. Laredo, Webb County, South Texas, USA, April
Four Mallard (Anas platyrhynchos) dabbling for food in icy water. The Netherlands, January
Zebra (Equus quagga) foal peering over its mother's striped back. Monochrome. Etosha National Park, Namibia, January
Hedge sparrow / Dunnock (Prunella modularis) nest with three eggs, nest exposed in hedge by summer hedge cutting, UK
Adult Fossa (Crytoprocta ferox) female portrait. Kirindy Forest, western Madagascar, November
Black and white ruffed lemur (Varecia variegata) hanging from tree, Madagascar, critically endangered
Pyramid of sea ice catching the first rays of the returning sun in the high Arctic, Svalbard, Spitsbergen, Norway, February 2009. Taken on location for the BBC series, Frozen Planet
A high view of mist shrouding an autumnal valley. Cadi Natural Park, Catalonia, Barcelona province, Pyrenees, Spain, October 2005
Close-up of scales of the Yellowbar Angelfish (Pomacanthus maculosus). Red Sea, Egypt
Stockdove (Columba oenas) shadow on the trunk of beech tree. Black Forest, Germany, April
Adult bull African Elephant (Loxononta africana) reaching up to feed on foliage on the banks of the Luangwa River, South Luangwa National Park, Zambia, October
Victorias riflebird (Ptiloris victoriae) male displaying in rainforest, Atherton Tablelands, Queensland, Australia, October
A female Spiny Seahorse (Hippocampus guttulatus) in weeds. Sardina, Gran Canaria, Canary Islands, East Atlantic Ocean, October
In-camera motion blur image of a female Yellow Seahorse (Hippocampus kuda). Lembeh Strait, Sulawesi, Indonesia, Molluca Sea, March
A batch of Leafy Seadragon (Phycodurus eques) eggs on the tail of a male. Wool Bay Jetty, Edithburgh, Yorke Peninsular, South Australia, November
Large / Pot Bellied Seahorse (Hippocampus abdominalis) face portrait. Blairgowrie Marina, Mornington Peninsular, Victoria, Australia, March
Silhouette of Red-fronted brown lemur (Eulemur rufus) leaping across gap in trees by moonlight, cathemeral behaviour, Ranomafana National Park, south east Madagascar, November
Asiatic lionesses and cubs (Panthera leo persica) drinking from pool, Gir Forest NP, Gujarat, India. Water is scarce in the Gir
Mute swans (Cygnus olor) newly hatched cygnets on pond, Yetholm Loch Scottish Wildlife Trust Reserve, Roxburghshire, Scotland, May
Highland bull (Bos taurus) face close up in freezing fog, Berwickshire Scotland, December
Goldfinches (Carduelis carduelis) squabbling near seed feeder, Berwickshire, Scotland, April
Brown trout (Salmo trutta) resting on river bed near the Aigas Field Studies Centre, Inverness-shire, Scotland, July
Clouded leopard (Neofelis nebulosa) Guwahati, Assam, India, captive
Mistle Thrush (Turdus viscivorus) in snow. Hertfordshire, England, UK, February
Southern plains grey langur ( Semnopithecus entellus / Presbytis entellus) close up face profile, Sawai Modhopur, Rajasthan, India
Boxer looking over closed door, standing on hind legs
Spotted scorpionfish (Scorpaena plumieri) lying motionless and camouflaged in soft corals. West Palm Beach, Gulf Stream, West Atlantic Ocean, Florida, USA
Banded jawfish (Opistognathus macrognathus) male incubating eggs in mouth, which are at a late stage of development and will soon be ready for release
