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Fallow deer (Dama dama) buck crossing road under street lights. London, UK. January
Wood sandpiper (Tringa glareola) at dusk silhouetted against setting sun, Karigasniemi, Northern Finland, May. Bookplate from Danny Greens The Long Journey North
Atlantic Salmon (Salmo salar) in pool on upstream spawning migration, silhouetted against sky. Quebec, Canada, August
Lion cub (Panthera leo) silhouetted at sunrise, taken on location for Pride tv series. September 2003
RF- Looking up under the gills of toadstools of Porcelain fungus (Oudemansiella mucida), Cornwall, UK
English oak tree (Quercus robur) at sunset, Monmouthshire Wales UK, March
Manta ray (Manta birostris), San Benedicto, Revillagigedo (Socorro) Islands, Mexico, East Pacific Ocean
Common Swallowtail butterfly (Papilio machaon) backlit showing wing structure. Podere Montecucco, Orvieto, Umbria, Italy, October
RF - Delicate grass and flowers silhouetted black against white background
Eagle owl (Bubo bubo) in flight through forest, backlit at dawn, Czech Republic, November. Captive
European Cave Spider (Meta menardi) in limestone cave. Plitvice Lakes National Park, Croatia. January
RF- Little blue heron (Egretta caerulea) and Great egret (Ardea alba) in lagoon at sunrise. Fort Myers beach, Gulf Coast, Florida, USA, March
Little blue heron (Egretta caerulea) in lagoon at sunrise, Fort Myers Beach, Gulf Coast, Florida, USA, March
Great white egret (Ardea alba) silhouetted at sunset, Lake Csaj, Pusztaszer, Hungary, February. Winner of the Portfolio category of the Terre Sauvage Nature Images Awards competition 2015
Hedge Parsley seed head (Torilis japonica) silhouetted at sunset, Norfolk, UK, November
Polar bear (Ursus maritimus) silhouetted at waters edge at sunset, Bernard Spit, off the 1002 Area, Arctic National Wildlife Refuge, North Slope, Alaska, USA, September. Vulnerable species
Common / Eurasian cranes (Grus grus) taking flight for roasting site, at sunset, silhouetted. Lake Hornborga, Sweden, April
Kelp gull (Larus dominicanus) on rocky shore with crashing waves at sunset, Bird Island, Lamberts Bay, Western Cape province, South Africa, September 2012
Two Great egrets (Ardea alba) wading, silhouetted at dawn, Keoladeo National Park, Bharatpur, Rajasthan, India
Masai giraffes (Giraffa camelopardalis tippelskirchi), at sunrise, Masai-Mara Game Reserve, Kenya. March
Straw-coloured fruit bats (Eidolon helvum) returning to daytime roost at sunrise. Kasanka National Park, Zambia
Stag beetle (Lucanus cervus) silhouetted against oak tree leaf. Elbe, Germany, June
Northern lights (Aurora borealis) glowing brightly over trees along Steese Highway, Cleary Point, outside Fairbanks, Alaska
Schooling Scalloped hammerhead sharks (Sphyrna lewini) Cocos Island National Park, Costa Rica
Cup fungus (Cookeina sp) Danum Valley, Sabah, Borneo
Black Grouse (Tetrao tetrix) displaying with breath vapor at dawn, Utajarvi, Finland, April
Brown-lipped snail (Cepaea nemoralis) on fungus, reproduced in a bokeh affect, Sheffield, England, UK. November. Commended in the Natures Studio Category of GDT competition 2014
African elephant (Loxodonta africana) herd silhouetted in distance walking through a storm, Masai-Mara Game Reserve, Kenya. September
Red deer (Cervus elaphus) stag bellowing during rut, Derbyshire, UK
Aerial view of two Gemsbok (Oryx gazella) in sand dunes, Namib Desert, Namibia
Deer mouse (Peromyscus maniculatus) on giant puffball mushroom, watching mosquito in the moonlight. Blackfoot Valley, Western Montana, USA, July
RF - Flock of Boat-tailed grackle (Quiscalus major) on wires at dawn. Crystal River, Florida, United States
Hoatzin (Opisthocomus hoazin) perched in tree, silhouetted at dusk, Cuyabeno, Sucumbios, Ecuador
Masai giraffe (Giraffa camelopardalis tippelskirchi) two silhouetted at sunrise, Msai Mara Game Reserve, Kenya
Kori bustard (Ardeotis kori) male, performing mating display, silhouetted at dusk, El Karama Wilderness Conservancy, northern Kenya
Tufted duck (Aythya fuligula) on water on a misty morning, Valkenhorst Nature Reserve, Valkenswaard, The Netherlands. June
Waterfowl at dawn with mist, including Great blue heron (Ardea herodias herodias), American avocet (Recurvirostra americana), Northern shoveler (Anas clypeata), Xochimilco wetlands, Mexico City
Silhouette of Red deer (Cervus elaphus) stag standing under a tree, Bradgate Park, Leicestershire, UK. October
Iberian lynx (lynx pardinus) in distance, silhouetted on horizon, Sierra de Andujar NP, Andalusia, Spain. December
Banded demoiselle dragonfly (Calopteryx splendens) male and female silhouetted, resting on reeds. Lower Tamar Lakes, Cornwall, England, UK. June
African elephant (Loxodonta africana) herd with calf, silhouetted, Botswana
Kudu (Tragelaphus strepsiceros) male silhouetted, Kruger National Park, South Africa. Digital silhouette
Silhouetted ferns in cloud-forest, Mashpi, Pichincha, Ecuador
Palm (Areaceae) leaves silhouetted in mist at dawn, Royal Chitwan National Park, Nepal
Pair of Red-eyed damselfly (Erythromma najas) perched on grass blade, at sunset, Poland. June
Loch Hope with hills silhouetted at sunset, Loch Hope, Sutherland, Scotland, November 2014
Ben Loyal from Kyle of Tongue in stormy light, Sutherland, Scotland, UK, December 2014
Roe deer (Capreolus capreolus) on a misty morning at the edge of a pond. Valkenhorst Nature Reserve, Valkenswaard, The Netherlands. December