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Red deer, (Cervus elaphus), stags silhouetted at sunset in winter, Scotland, UK. FebruaryRed deer, (Cervus elaphus), stags silhouetted at sunset in winter, Scotland, UK.February
An Orange lacewing butterfly (Cethosia penthesilea), Cairns Botanical Gardens, Queensland, Australia. Captive. August
Green-veined white butterfly (Artogeia / Pieris napi) resting on reed in late evening light, Volehouse Moor, Devon, UK. July
Abstract patterns of reflected sky and clouds on the ocean with a silhouetted Red-faced cormorant (Phalarocorax urile) in flight, St. Paul Island, Pribilof Islands, Alaska, USA, July
Great Egret (Ardea alba), about to land in water, backlit, Bolsa Chica Ecological Reserve, California, USA, October
Mixed Blackbird flock, mostly Red-winged Blackbirds (Agelaius phoeniceus), in flight against backdrop of autumn foliage in November, Bosque Del Apache National Wildlife Refuge, New Mexico, USA
European otter (Lutra lutra) group with one shaking off water, Kiskunsagi National Park, Hungary, October
European otter (Lutra lutra) shaking off water, Kiskunsagi National Park, Hungary, January
Wood warbler (Phylloscopus sibilatrix) singing whilst perched on branch, silhouetted at dawn. Sheffield, England, UK. May
Red fox (Vulpes vulpes) silhouetted, Sado Estuary, Portugal. September
Synchronous fire flies (Pteroptyx sp. ) mangroves along the Kinabatangan River, Sabah, BorneoSynchronous fire flies (Pteroptyx sp.) mangroves along the Kinabatangan River, Sabah, Borneo
Fallow deer (Dama dama) at night, Gyulaj, Hungary Third place in the Nature Portfolio category of the World Press Photo Awards 2017
European Greenfinch (Chloris chloris) flying at dawn. When backlit the tiny transparent barbs that make up the feather act like a prism revealing vivid colours in the early light
Sumatran orangutan (Pongo abelii) female moving between lianas, Gunung Leuser National Park, UNESCO World Heritage site, November
Aerial view of two Gemsbok (Oryx gazella) in sand dunes, Namib Desert, Namibia
Grey heron (Ardea cinerea) catching fish in lake, Pusztaszer, Hungary, April
Killer whale (Orcinus orca) adult male surfacing at dusk surrounded by birds, who watch for herrings (Clupea harengus) left after they have fed, Andenes, Andoya island, North Atlantic Ocean, Norway
Praying mantis nymph silhouetted through leaf, . Raja Ampat, Western Papua, Indonesian New Guinea
Chinese lanterns (Physalis alkekengi) skeletons, silhouette on white background
Hedge parsley (Torilis japonica) close up pattern of dead seed heads. Silhouette on white background
Chinese Lanterns (Physalis alkekengi), Hydrangea flowers, Poplar leaves (Populus) and Honesty skeletons (Lunaria annua), silhouettes on white background
Stonehenge at sunrise, Salisbury Plain, Wiltshire, England, UK, June 2006
Corfe Castle in morning mist, Corfe, Isle of Purbeck, Dorset, England, UK. September 2013
Bottlenosed dolphin (Tursiops truncatus) porpoising during annual sardine run, Port St Johns, South Africa. June
Angolan giraffes (Giraffa camelopardalis angolensis) and black rhinoceros (Diceros bicornis) reflecting in waterhole at sunset, Okaukuejo, Etosha National Park, Namibia, Africa
Nilgai or Blue bull (Boselaphus tragocamelus), silhouette of male at sunset, with Black drongo bird on its back. Velavadar National Park, Gujarat, India
Humayuns night frog (nyctibatrachus humayuni), male guarding cluster of eggs on leaf. Western Ghats Amboli, Maharashtra, India. Endemic
Mountain landscape at Tangjiahe National Nature Reserve, Sichuan Province, China April, 2015
Spoonbill (Platalea leucorodia) carrying nest material, The Netherlands
Ripe Barley (Hordeum vulgare) ears in field ready for harvest and the rising sun, Stone Henge, Salisbury, England, UK, August
Borage flower bud (Borago officinalis) near Nice, south of France, June
Peruvian mantis (Oxyopsis peruviana) silhouette with sun behind, captive, occurs in South America
Groot Schietveld at sunrise, Wuustwezel, Belgium, May 2010
Scots pine tree (Pinus sylvestris) at sunrise, Klein Schietveld, Brasschaat, Belgium. September
Scots pine trees (Pinus sylvestris) on island in wetlands, Klein Schietveld, Brasschaat, Belgium
Boababs (Adansonia grandidieri) silhouetted at dawn, Allee des Baobabs / Avenue of the Baobabs, Morondave, Madagascar
RF - Reticulated Glass Frog (Hyalinobatrachium valerioi) backlit showing highly translucent body. Osa Peninsula, Costa Rica
Pheasant (Phasianus colchicus) male flying to roost at sunset with trees silhouetted in background England, UK. December
Dingo (Canis lupus dingo) fighting on a beach. Fraser Island UNESCO World Heritage Site. Queensland, Australia, November
RF - Grizzly Bear (Ursus arctos) at dawn, Lake Clarke National Park, Alaska, September
RF- White storks (Ciconia ciconia) nesting on poles erected by the city of Caceres in Extremadura, Spain, to compensate for the removal of an old farmhouse supporting a colony of storks
Frog spawn {Rana temporaria} just a 1-2 days old, backlit on lightbox, Cornwall, UK. January 2012
Black kitten in garden, Germany
Mute swan (Cygnus olor) at sunrise silhouetted in waters of Lamlash Bay, Isle of Arran, South Arran Marine Protected Area, Scotland, UK, August
Sea stack and Talisker Beach, Isle of Skye, Inner Hebrides, Scotland, UK, September 2013
View from Isle of Skye across Sound of Rsay to Rona and Torridon Hills, at dawn, Inner Hebrides, Scotland, UK, April 2014
Masai giraffe (Giraffa camelopardalis tippelskirchi) herd walking, Masai-Mara Game Reserve, Kenya. October
Fallow deer (Dama dama) buck crossing road under street lights. London, UK. January
