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Grey heron (Ardea cinerea) in rain with another heron behind, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Red-legged honeycreeper (Cyanerpes cyaneus) two males on barbed wire. Costa Rica
Two Buzzards (Buteo buteo) fighting, Lake Csaj, Kiskunsagi National Park, Hungary. February
Cormorants (Phalacrocorax carbo) squabbling over fish, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Two Burrowing owls (Athene cunicularia) fledgling with adult about to take off, Pantanal, Brazil
Corncrake (Crex Crex) calling, with breath vapor in early morning, Kiskunsagi National Park, Hungary
Montezuma oropendola (Psarocolius montezuma) hanging upside down as part of a courtship display. Costa Rica
Synchronous fire flies (Pteroptyx sp. ) mangroves along the Kinabatangan River, Sabah, BorneoSynchronous fire flies (Pteroptyx sp.) mangroves along the Kinabatangan River, Sabah, Borneo
Grey herons (Ardea cinerea) fighting, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Grey heron (Ardea cinerea) squabbling, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. February
Flock of Tree sparrows (Passer montanus) in flight, with a Greenfinch (Chloris chloris) blurred motion, Pacsmag, Hungary
Double-crested basilisk (Basiliscus plumifrons) running across water surface, Santa Rita, Costa Rica
Shadowfin soldierfish (Myripristis adusta) West Papua, Indonesia
Montane horned frog (Megophrys kobayashii) in the leaf litter at night. Mount Kinabalu (around 1400m asl), Sabah, Borneo
Shrimpfish / Razorfish (Aeoliscus punctatus / strigatus) have adapted to a vertical posture and swim with their head downward. Lembeh, North Sulawesi, Indonesia
Wide angle view of Demoiselle cranes (Anthropoides virgo) landing in chugga ghar (bird feeding enclosure), Khichan, Western Rajasthan, India. December
Demoiselle crane (Anthropoides virgo) flock taking flight during winter migration. Khichan, Western Rajasthan, India. February 2015
Red-and-green macaws (Ara chloropterus) perched and in flight over Buraco das Araras, the Sinkhole of the Macaws, Jardim, Mato Grosso do Sul, Brazil. September
Early morning shafts of light through deciduous forest. Bandhavgarh National Park, India
Sun beam at sun rise over the Grand Canyon of the Yellowstone, Yelllowstone National Park, Wyoming, USA. January 2016
Anthers in garden, Vilcabamba, Ecuador. November
Cluster of Burnt or Burnt-tip Orchids (Neotinea ustulata) in a Alpine meadow. Tyrol, Austria
Aerial view of Belas Knap, a neolithic chambered long barrow on the Cotswold Way, Winchcombe, Gloucestershire, UK. Shot by aerial drone with Ca permit. January 2016
Nurse shark (Ginglymostoma cirratum) and Horse-eye jacks (Caranx latus) Shark Ray Alley, Hol Chan Marine Reserve, Belize Barrier Reef, Belize
Marsh fritillary butterfly (Euphydryas aurinia) feeding on Wild garlic / Ramsons (Allium ursinum) at Strawberry Banks, Gloucestershire Wildlife Trust (GWT), Nature Reserve, Gloucestershire, UK. May
Whale Shark (Rhincodon typus) and Golden trevally (Gnathanodon speciosus) Cenderawasih Bay, West Papua, Indonesia
Jack snipe (Lymnocryptes minima) feeding in margins of watercress bed, Hertfordshire, England, UK, January
Whale shark (Rhincodon typus) front view portrait, Cenderawasih Bay, West Papua. Indonesia
Aerial view of African elephants (Loxodonta africana) looking for the shade under trees on a very hot day, Northern Botswana
Whale shark (Rhincodon typus) near surface, viewed from above, Cenderawasih Bay, West Papua. Indonesia
Southern Lechwe (Kobus lechwe) leaping through water, Okavango Delta, Botswana. Low angle remote camera shot
Marsh helleborine (Epipactis palustris) close up of two flowers showing front view, Hertfordshire, England, UK, June. Focus stacked image
Two coloured mason bee (Osmia bicolor) bee that nests in old snail shells and then carries bits of twig or vegetation to cover the shell, Buckinghamshire, England, UK, June
Gray whale (Eschrichtius robustus) aerial view of blowing during whale migration, Coast of California, USA, Eastern Pacific Ocean
Black-backed jackals (Canis mesomelas) two fighting with Plains / Burchell zebra (Equus burchelli) in foreground, Kalahari Desert, Nxai Pans National Park. Botswana
Killer whale (Orcinus orca) hunting for herring fish (Clupea harengus), Andenes, Andoya island, North Atlantic Ocean, Norway, April
Reef manta (Manta alfredi) mouth open feeding at night, South Ari atoll, Maldives, Indian Ocean
Copper-rumped hummingbird (Saucerottia tobaci) feeding, Tobago, West Indies
Killer whale (Orcinus orca) hunting inside herring baitball (Clupea harengus), Andenes, Andoya island, North Atlantic Ocean, Norway, April
Rainbow scarab (Phanaeus vindex) male on white background. Florida, USA. Controlled conditions
Plain / Burchells zebras in dust (Equus burchelli) Nxai Pan National Park, Botswana
Cockroaches (Periplaneta americana) on wall of Gomantong Cave. Sabah, Borneo. Malaysia
Spider webs covered in dew, on a flowering Gorse bush, Peak District National Park, UK October
Historical illustration of Common octopus (Octopus vulgaris) Pierre Denys de Montfort engraving from Histoire Naturelle Generale et Particuliere des Mollusques, 1801-1802
Giant honey bee nest (Apis dorsata) up in a giant Mengaris tree (Koompassia excelsa) dipterocarp forest in Danum Valley, Sabah, Borneo, Malaysia
Leopard (Panthera pardus) walking along waterhole, reflected in the water at dusk. Londolozi Private Game Reserve, Sabi Sand Game Reserve, South Africa
Spirit / Kermode bear (Ursus americanus kermodei) catching fish in river, most likely salmon, the Great Bear Rainforest, British Columbia, Canada, October
Coastal brown bears (Ursus arctos) fighting, Lake Clarke National Park, Alaska, September