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Upland goose (Chloephaga picta) goslings, Sea Lion Island, Falkland Islands, October
King penguin (Aptenodytes patagonicus) chick creche, Volunteer Point, East Falkland, Falkland Islands, October
Striated caracara (Phalcoboenus australis) group with one landing from above with only wings and tail visible. Carcass Island, Falkland Islands, October
King penguin (Aptenodytes patagonicus) colony, Volunteer Point, East Falkland, Falkland Islands, October
Aerial view of fish farm, La Guittiere Marsh, South Vendee, France, July 2017
Mute swan (Cygnus olor) cygnets with head submerged while feeding underwater, with mother watching and helping, Derbyshire, England, UK. June
Tree and rock formations at Brimham Rocks, Harrogate, Yorkshire. Comprised of Carboniferous age, Millstone Grit, these strange rock formations form by variable erosion of soft
Leaves of a Common aspen tree (Populus tremula) on ground in autumn, Aragon, Spain, November
Willow (Salix) plantation, Cuenca, Castilla-La Mancha, Spain, December
Footpath sign in Norfolk countryside with Blackthorn hedgerow (Prunus spinosa) in flower. Norfolk, England, UK. April
Fallow deer (Dama dama) at night, Gyulaj, Hungary Third place in the Nature Portfolio category of the World Press Photo Awards 2017
Eurasian blackcaps (Sylvia atricapilla) caught on limed stick, Cyprus. Finalist in The Wildlife Photojournalist Award: Single Image Category of the Wildlife Photographer of the Year Awards (WPOY)
African lion (Panthera leo) at waterhole at night, Mkuze, South Africa Third place in the Nature Portfolio category of the World Press Photo Awards 2017
Zebra at waterhole at night, Mkuze, South Africa Third place in the Nature Portfolio category of the World Press Photo Awards 2017
Ordesa y Monte Perdido National Park, Huesca, Aragon, Spain, July 2016
Northern Lights (Aurora Borealis) over Hornafjordur, eastern Iceland, February 2016
Bornean orangutan (Pongo pygmaeus) male, Gunung Palung National Park, Borneo
Eurasian griffon vulture (Gyps fulvus) in flight, Cevennes, France, March 2016
Carp (Cyprinus carpio) in oxbow lake of the Aare River, Switzerland
Polar bear (Ursus maritimus) on an iceberg, Baffin Bay, Canada. September
Red leaf monkey (Presbytis rubicunda) female feeding in Strangler fig tree (Ficus dubia) Gunung Palung National Park, Kalimantan, Borneo, Indonesia
Wild boar (Sus scrofa) running across snow covered field, Vosges, France, January
European robin (Erithacus rubecula), Vosges, France, December
Hawksbill turtle (Eretmochelys imbricata) cruising along the drop off of a coral reef. Bloody Bay Wall, Little Cayman, Cayman Islands. British West Indies. Caribbean Sea
Blue morpho (Morpho helenor) butterflies feeding on mango fruit, butterfly observatory, La Paz Waterfall Gardens, Costa Rica
California sealion, (Zalophus californianus), playing with seastar, Los Islotes, Sea of Cortez, Baja California, Mexico, East Pacific Ocean
Ring tailed lemur (Lemur catta) backlit in late afternoon light, Berenty Private Reserve, southern Madagascar
Spotted Lake, a saline endorheic alkali lake, a medicine lake for the Okanagan Syilx people, British Columbia, Canada, July
Leaf mimic bush cricket or katydid (unknown species, family Tettigoniidae) in the middle of moulting its skin. Moulting from one sub-adult stage (instar) to another is known as ecdysis
Giant Manta Ray (Manta birostris), with Remoras (Echeneididae) Socorro Island, Revillagigedo Archipelago Biosphere Reserve (Socorro Islands), Pacific Ocean, Western Mexico, March
White fronted capuchin monkey (Cebus capucinus) feeding, March, Costa Rica
Madagascar / Malagasy kingfisher (Alcedo vintsioides) perching, Anjajavy Private Reserve, north west Madagascar
Afrian lioness (Panthera leo) playing with her juvenile cub aged 2 years in Duba Plains concession. Okavango delta, Botswana
Male Indri (Indri indri) leaping through the rain forest canopy. Andasibe-Mantadia National Park, eastern Madagascar. Endangered
Wicken Fen wetlands with windmill water pump and Phragmites reeds, Cambridgeshire, England, May
Cultivated Crab apple tree (Malus-x-robusta Red-Sentinel ) blossom
Landscape of the Virgin Komi Forests UNESCO World Heritage site at sunrise. These are the largest virgin forests in Europe. Ural Mountains, Komi Republic, Russia. August 2016
River through Virgin Komi Forests UNESCO World Heritage site, the largest virgin forests in Europe. Ural Mountains, Komi Republic, Russia. August 2016
Puma (Puma concolor) looking upwards, Corcovado National Park, Costa Rica, May
Meerkats (Suricata suricatta), Kgalagadi Transfrontier Park, Northern Cape, South Africa, January
Lioness (Panthera leo) drinking at night, Zimanga Private Game Reserve, KwaZulu-Natal, South Africa
White rhino (Ceratotherium simum) at waterhole at night, Zimanga Private Game Reserve, KwaZulu-Natal, South Africa
Young ground squirrels (Xerus inauris) fighting, Kgalagadi Transfrontier Park, Northern Cape, South Africa, January
Impala (Aepyceros melampus) with redbilled oxpecker (Buphagus erythrorhynchus), Kruger National Park, South Africa
European Greenfinch (Chloris chloris) flying at dawn. When backlit the tiny transparent barbs that make up the feather act like a prism revealing vivid colours in the early light
English oak tree (Quercus robur) in field of Oilseed rape / rapeseed (Brassica napus), UK, May 2017
Pied crows (Corvus albus) perched on horns of antelope skull. Zimanga Private Game Reserve, KwaZulu-Natal, South Africa
Adult King Vulture (Sarcoramphus papa) Laguna del Lagarto, Boca Tapada, Caribbean slope, Costa Rica, Central America