Heather Angel Collection (page 4)

Scarab beetle (Scarabaeoidea) pair mating inside Tulip (Tulipa micheliana) flower. Beetles covered in pollen. Uzbekistan, April

Large-flowered evening primrose (Oenothera glazioviana). Bulls eye in centre of flower fluorescing in UV light guides pollinators to nectaries. At night in garden. Surrey, England, UK

Sun bromeliad (Fascicularia bicolor). Central leaves turn red to attract hummingbird pollinators before flowers open. Cultivated in garden. Surrey, England, UK

Spider flower (Cleome sp) with long gonophores. View from above. Cultivated in garden, Surrey, England, UK. Native to South America

Silver Y moth (Autographa gamma) nectaring on Evening primrose (Oenothera glazioviana) with proboscis extended. At night in garden. Surrey, England, UK. August

Malachite sunbird (Nectarinia famosa) male nectaring on King protea (Protea cynaroides). Cape Province, South Africa. August

Wasp (Vespula germanica) nectaring on Pineapple lily (Eucomis comosa). Pollen is deposited on underside of wasps hairy body and transferred to stigma of next flower visited

Honey bee (Apis mellifera) covered in pollen emerging from nectaring in Crocus (Crocus sp). On nature reserve, Surrey, England, UK. March

Honey bee (Apis mellifera) with pollen grains on back. Inside male Squash (Cucurbita sp) flower. Cultivated in garden, Surrey, England, UK. September

Wool carder bee (Anthidium manicatum) female entering Rusty foxglove (Digitalis ferruginea) flower. Pollen transferring to head from stamens on upper lip of flower

Common bee fly (Bombylius major) nectaring on Primrose (Primula vulgaris). This insect is the main pollinator of Primrose. In nature reserve, Surrey, England, UK, April

Bumblebee scarab (Amphicoma vulpes) pair mating in Tulip (Tulipa armena) flower. Pollen picked up on beetles and transferred to other flowers. Near Karabel Pass, Turkey. May

Hazel (Corylus avellana) catkins dispersing pollen with female flower above on branch. Controlled condition.s Surrey, England, UK

Musquia (Musschia aurea) flower. Cultivated in glasshouse, UK. Pollinated by Lizard (Lacerta dugesii) and Madeira bumblebee (Bombus maderensis) in Madeira

Texas bluebonnets (Lupinus texensis) and Indian paintbrush (Castilleja coccinea) amongst wildflowers in ditch beside byway. Rains triggered germinaton of annual seeds. Cuero, Texas, USA. April

Orange breasted sunbird (Anthobaphes violacea), male nectaring on Heather (Erica sp). Kirstenbosch Botanic Garden, Cape Town, South Africa. February

Purple throated mountain gem hummingbird (Lampornis calolaemus) hovering as it nectars on Porterweed (Stachytarpheta frantzii). Costa Rica

Yellow mariposa lily (Calochortus luteus) with dark nectar guides. Cultivated in glasshouse, Surrey, England, UK. Endemic to California

Winter aconite (Eranthis hymenalis) in UV light, pollen on outer stamens that have dehisced fluorescing. Focus stacked

Common rue (Ruta graveolens) flower in visible light, nectar at base of petals. Stamens flip up sequentially with one still folded over the receptacle. Surrey, England, UK

Common rue (Ruta graveolens) flower in UV light, pollen and nectar at base of petals fluorescing. Insect pollinators are able to see position of rewards on offer. Surrey, England, UK

Tulip tree (Liriodendron tulipifera) flower with orange band to attract bees, produces copious nectar. In visible light. Surrey, England, UK. Native to USA. Focus stacked. Sequence 1/2

Tulip tree (Liriodendron tulipifera) flower, pollen and nectar fluorescing in UV light. Surrey, England, UK. Native to USA. Focus stacked. Sequence 2/2

Two-coloured pineapple lily (Eucomis bicolor) in visible light. Cultivated in garden, Surrey, England, UK. Native to South Africa. Focus stacked. Sequence 1/2

Porcelain orchid (Chloraea magellanica) with osmophores on lip fluorescing in UV light. Chile. Controlled conditions, focus stacked. Sequence 2/2

Foxglove (Digitalis purpurea) corollas with spatter mark nectar guides. Pollinated by long tongued bumblebees able to reach nectar

Seville orange (Citrus u aurantium) flower. Cultivated in glasshouse, Surrey, England, UK
Seville orange (Citrus × aurantium) flower. Cultivated in glasshouse, Surrey, England, UK. Focus stacked. Sequence 1/2

Seville orange (Citrus u aurantium) flower fluorescing in UV light, colour attractive
Seville orange (Citrus × aurantium) flower fluorescing in UV light, colour attractive to pollinators. Cultivated in glasshouse, Surrey, England, UK. Focus stacked. Sequence 2/2

Sacred lotus (Nelumbo nucifera) in visible light. Cultivated in glasshouse, Surrey, England, UK. Sequence 1/2

Southern catalpa (Catalpa bignonioides) flowers. Recently opened with yellow nectar guides, purple in older pollinated flowers. Surrey, England, UK. Native to USA

Wasp nectaring on Marsh marigold (Caltha palustris), picking up pollen on head, thorax and hairs on body. Turkey. May

Flame lily (Gloriosa superba), timelapse sequence from opening bud to flowering, tepals becoming reflexed. Flower pollinated by butterflies. Controlled conditions

Narcissus fly (Merodon equestris) male nectaring on Rock rose (Cistus x hybridus) in garden, Surrey, England, UK

Garden bumblebee (Bombus hortorum) queen nectaring on Comfrey (Symphytum Hidcote Pink ) flowers. In garden, Surrey, England, UK. March

Red admiral (Vanessa atalanta) basking in sun, on Snowdrops (Galanthus nivalis). In garden, Surrey, England, UK. February

Buff tailed bumblebee (Bombus terrestris) queen nectaring on Rhododendron (Rhododendron sp), pollen from anthers on backside and legs. Surrey, England, UK. March

Red-legged bumble-moth (Homoeocera gigantea) on Lichen covered tree trunk. Costa Rica

Great yellow bumblebee (Bombus distinguendus) nectaring on Kidney vetch (Anthyllis vulneraria). RSPB Balranald Nature Reserve, North Uist; Outer Hebrides, Scotland, UK. July

Rose chafer (Cetonia aurata) feeding on Rock rose (Cistus x hybridus) pollen. Cuts in petals where chafers legs were inserted to prevent it from falling off. In garden, Surrey, England, UK. June

Marmalade hoverfly (Episyrphus balteatus) feeding on Golden apple (Lilium carniolicum) pollen, transfers pollen on body to other flowers. In high grassland, Mount Vournos, northern Greece

White-collar burnet moth (Zygaena carniolica) nectaring on Catnip (Nepeta nuda). Greece. July

Bee beetle (Trichius fasciatus), a bee mimic, nectaring on Umbel-flowered catchfly (Silene compacta). Picks up pollen on hairy body. Russian Caucasus. June

Honeybee (Apis mellifera) with full pollen baskets and pollen on head, thorax and abdomen. Bee withdrawing from Snowdrop (Galanthus nivalis) flower after nectaring. In garden, Surrey, England, UK

Marsh fritillary (Euphydryas aurinia) resting on Caucasian inula (Inula orientalis) following light rain. Russian Caucasus. June

Carpenter bee (Xylocopa caffra) male robbing nectar from Ninepin heath (Erica mammosa), not pollinating flower. The Cape, South Africa. February

Scarab beetles sleeping overnight in Persian marsh orchid (Dactylorhiza umbrosa). Turkey. June

Red mucronata (Balanophora harlandii), a parasite on woody roots with flowers resembling fungi. Wild Elephant Valley, Xishuangbanna, Yunnan, China

Pere Davids deer (Elaphurus davidianus) two stags boxing amongst herd of hinds. Captive bred in reserve after reintroduction from Woburn Abbey, England, UK

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