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Grey wolf (Canis lupus) pack resting on a snow ledge, Hayden Valley, Yellowstone National Park, USA. January
Red fox (Vulpes vulpes) resting in snow, Silver Gate, Yellowstone National Park, USA. January
American pine marten (Martes americana) standing on hind legs in deep snow, Yellowstone National Park, USA. January
Female Bison (Bison bison) with young calf, walking over snow in front of frost-covered tree, Yellowstone National Park, USA. February
Bison (Bison bison) being circled by wolves (Canis lupus) hunting in snow, Yellowstone National Park, USA. January
Wolf (Canis lupus) standing in snow, Yellowstone National Park, USA. January
Four Wolves (Canis lupus) walking in snow, Yellowstone National Park, USA. January
Wolf (Canis lupus) walking in snow, Yellowstone National Park, USA. January
Two Wolves (Canis lupus) standing in deep winter snow, Yellowstone National Park, USA. January
Red fox (Vulpes vulpes) in its winter coat, Yellowstone National Park, Wyoming, USA. October
Great grey owl (Strix nebulosa) preparing to land, in tree, Yellowstone National Park, Wyoming, USA
Coyote (Canis latran) in sagebrush grassland, Yellowstone National Park, Wyoming, USA
Grand Prismatic Springs on cold winter day with mist / vapour, Yellowstone National Park, USA, January
North American river otter (Lontra canadensis) on snow covered bank, reflected in icy water. Yellowstone National Park, USA, January
RF - North American river otters (Lontra canadensis) (probably female with two juveniles) on the frozen river edge.Yellowstone National Park, USA. January. Digitally stitched image
RF - Red fox (Vulpes vulpes) on the edge of coniferous forest. Yellowstone National Park, USA. January (This image may be licensed either as rights managed or royalty free.)
RF - Lodgepole pines (Pinus contorta) covered in heavy early morning hoar frost. Yellowstone National Park, USA. January. Digitally stitched image
Red fox (Vulpes vulpes) walking through deep winter snow. Hayden Valley, Yellowstone, USA. January
North American river otter (Lontra canadensis) female and cub walking across snow, reflected in water. Yellowstone National Park, USA, January
Coyote (Canis latrans) standing in snow, dwarfed by frosty trees. Yellowstone National Park, USA, January
RF - Petrified trees and travetine cascade. Geothermal feature at Mammoth Hot Springs. Yellowstone National Park, USA. (This image may be licensed either as rights managed or royalty free.)
RF - Red fox (Vulpes vulpes) in deep snow. Yellowstone National Park, USA. January (This image may be licensed either as rights managed or royalty free.)
RF - North American river otters (Lutra canadiensis) on the frozen river edge. Yellowstone National Park, USA. January (This image may be licensed either as rights managed or royalty free.)
Coyote (Canis latrans) foraging in deep winter snow disturbed by grazing American bison (Bison bison) Hayden Valley, Yellowstone National Park, Wyoming, USA. January
North American river otters (Lutra canadiensis) probaly female with two juveniles, on the frozen river edge. Upper Yellowstone River, Hayden Valley, Yellowstone, USA. January
North American river otter (Lutra canadiensis) two on frozen river edge. Upper Yellowstone River, Hayden Valley, Yellowstone National Park, Wyoming, USA. January
Coyote (Canis latrans) foraging in snow. Hayden Valley, Yellowstone National Park, USA. January 2019
American Bison (Bison bison) frost covered male, standing by dead trees. Firehole River Valley. Yellowstone National Park, Wyoming, USA. January
Rocky mountain bighorn sheep (Ovis canadensis canadensis) grazing. Lamar Valley, Yellowstone National Park, Wyoming, USA. January
RF - Herd of American Bison (Bison bison) around geo-thermal features. Firehole River Valley. Yellowstone National Park, Wyoming, USA. January 2008
Sun beam at sun rise over the Grand Canyon of the Yellowstone, Yelllowstone National Park, Wyoming, USA. January 2016
Grizzly bear (Ursus arctos horribilis) female with sub-adult cub rolling on back, in grassland. Yellowstone National Park, Wyoming, USA. June
RF - Red fox (Vulpes vulpes) foraging in snow covered valley. Hayden Valley, Yellowstone National Park, USA. February 2019. (This image may be licensed either as rights managed or royalty free.)
Red fox (Vulpes vulpes) sequence of snow diving whilst hunting for rodents. Hayden Valley, Yellowstone National Park, USA. February 2019. Digital composite
North American river otter (Lutra canadiensis), two standing at edge of frozen Upper Yellowstone River, Hayden Valley, Yellowstone, USA. February
Steam rising from Grand Prismatic thermal pool in snow covered landscape. Midway Geyser Basin, Yellowstone National Park, USA. February 2019
Coyote (Canis latrans) standing near waters edge, in snow. Hayden Valley, Yellowstone National Park, USA. January 2019
Conifer sapling emerging from snow drift with cornice. Hayden Valley, Yellowstone National Park, Wyoming, USA. January
Wolf (Canis lupus) lone male walking in shadow of the Teton Mountains. Snake River valley near Jackson Hole. Grand Teton National Park, Wyoming, USA. January
Coyote (Canis latrans) jumping in snow with its prey, Yellowstone National Park, Wyoming, USA, February
North American Bison (Bison bison) coated in frost standing on snow, Yellowstone National Park, Wyoming, USA, January
American elk (Cervus elaphus canadensis) stag sticking tongue out, Yellowstone National Park, Wyoming, USA
American barn swallow (Hirundo rustica erythrogaster) in flight catching insects over the Gallatin River, Yellowstone National Park, Montana, August
Coyote (Canis latrans) in snow, Yellowstone National Park, Wyoming, USA, February
RF - Coyote (Canis latrans) in snow, Yellowstone. February
Red fox (Vulpes vulpes) resting and sitting on snow, Yellowstone National Park, USA, February
Black bear (Ursus americanus) mother with cub. Yellowstone National Park, Wyoming, USA, May
