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North American Bison (Bison bison) coated in frost standing on snow, Yellowstone National Park, Wyoming, USA, January
RF - Bison (Bison bison) breathing in the cold air, Yellowstone National Park, USA, February
American bison (Bison bison) female covered in hoar frost near hot spring, portrait. Midway Geyser Basin, Yellowstone National Park, USA. February
Herd of American bison (Bison bison) in snow, Yellowstone National Park, Wyoming, Yellowstone, January
Bobcat (Lynx rufus) standing on branch in snow. Madison River Valley, Yellowstone National Park, Wyoming, USA. January
Bison (Bison bison) herd walking in snow, Yellowstone National Park, Wyoming, USA, February
American bison (Bison bison) male walking through deep snow. Hayden Valley. Yellowstone National Park, Wyoming, USA. January
Bison (Bison bison) in snow. Yellowstone National Park, USA, February
Bison (Bison bison) female with calf walking through snow in front of frost-covered forest, Yellowstone National Park, USA
Female American buffalo (Bison bison) walking across grassland with newborn calf, Yellowstone National Park, Wyoming, USA. May
Coyote (Canis latrans) pouncing, hunting technique in Yellowstone National Park, Wyoming, USA
Yellowstone River. Yellowstone National Park, Wyoming, USA. September
Elk (Cervus canadensis) bull and cw sniffing noses, Yellowstone National Park, Wyoming, USA. September
Red fox (Vulpes vulpes) running through snow. Hayden Valley, Yellowstone, USA. January
Red fox (Vulpes vulpes) adult hunting for rodents by snow diving in deep snow. Hayden Valley, Yellowstone, USA. January
Red fox (Vulpes vulpes) in mid air, snow diving / pouncing whilst hunting for rodents. Hayden Valley, Yellowstone National Park, USA. February
Pack of wolves (Canis lupus). Yellowstone National Park, Wyoming, USA. January
Male American Bison (Bison bison) walking through snow. Firehole River Valley, Yellowstone National Park, Wyoming, USA. January
Trumpeter swans (Cygnus buccinator) on the edge of the Upper Yellowstone River. Hayden Valley, Yellowstone National Park, Wyoming, USA. January
Coyote (Canis latrans) hunting after a fresh snowfall, Yellowstone National Park, Wyoming, USA. October
Grizzly bear (Ursus arctos horribilis) running along shore of Yellowstone Lake at sunrise, Yellowstone National Park, Wyoming, USA
North American river otter (Lontra canadensis) female and cub walking across snow, reflected in water. Yellowstone National Park, USA, January
North American river otter (Lutra canadiensis) two on frozen river edge. Upper Yellowstone River, Hayden Valley, Yellowstone National Park, Wyoming, USA. January
RF - American bison (Bison bison) male standing near Punch Bowl thermal pool, green algae in foreground. Yellowstone National Park, USA
Coyote (Canis latrans) foraging in snow. Hayden Valley, Yellowstone National Park, USA. January 2019
Rocky mountain bighorn sheep (Ovis canadensis canadensis) grazing. Lamar Valley, Yellowstone National Park, Wyoming, USA. January
North American Bobcat (Lynx rufus) stalking along the edge of the Madison River. Yellowstone National Park, Wyoming, USA. January
Black bear (Ursus americanus), two cubs playing, climbing up tree trunk. Yellowstone National Park, Wyoming, USA. June
Red fox (Vulpes vulpes) sequence of snow diving whilst hunting for rodents. Hayden Valley, Yellowstone National Park, USA. February 2019. Digital composite
North American river otter (Lutra canadiensis), two standing at edge of frozen Upper Yellowstone River, Hayden Valley, Yellowstone, USA. February
American bison (Bison bison) female grazing near thermal pool, snow on ground. Biscuit Geyser Basin, Yellowstone National Park, USA. February 2019
American bison (Bison bison), two females covered in hoar frost near hot spring, portrait. Midway Geyser Basin, Yellowstone National Park, USA. February
Morning Glory thermal pool surrounded by snow, coniferous forest in background. Near Daisy Geyser Basin, Yellowstone National Park, USA. January 2019
Conifer sapling emerging from snow drift with cornice. Hayden Valley, Yellowstone National Park, Wyoming, USA. January
RF - American Bison (Bison bison) male in snow covered in frost, standing by dead trees. Firehole River Valley. Yellowstone National Park, Wyoming, USA. January
Rainclouds over Madison Valley, Beaverhead National Forest, Greater Yellowstone Ecosystem, Madision Mountains, Montana USA, July
Grey wolf (Canis lupus) pack resting on a snow ledge, Hayden Valley, Yellowstone National Park, USA. January
Red fox (Vulpes vulpes) resting in snow, Silver Gate, Yellowstone National Park, USA. January
American pine marten (Martes americana) standing on hind legs in deep snow, Yellowstone National Park, USA. January
Female Bison (Bison bison) with young calf, walking over snow in front of frost-covered tree, Yellowstone National Park, USA. February
Bison (Bison bison) being circled by wolves (Canis lupus) hunting in snow, Yellowstone National Park, USA. January
Wolf (Canis lupus) standing in snow, Yellowstone National Park, USA. January
Four Wolves (Canis lupus) walking in snow, Yellowstone National Park, USA. January
Wolf (Canis lupus) walking in snow, Yellowstone National Park, USA. January
Two Wolves (Canis lupus) standing in deep winter snow, Yellowstone National Park, USA. January
Red fox (Vulpes vulpes) in its winter coat, Yellowstone National Park, Wyoming, USA. October
Great grey owl (Strix nebulosa) preparing to land, in tree, Yellowstone National Park, Wyoming, USA
Coyote (Canis latran) in sagebrush grassland, Yellowstone National Park, Wyoming, USA
Grand Prismatic Springs on cold winter day with mist / vapour, Yellowstone National Park, USA, January
North American river otter (Lontra canadensis) on snow covered bank, reflected in icy water. Yellowstone National Park, USA, January
RF - North American river otters (Lontra canadensis) (probably female with two juveniles) on the frozen river edge.Yellowstone National Park, USA. January. Digitally stitched image
RF - Red fox (Vulpes vulpes) on the edge of coniferous forest. Yellowstone National Park, USA. January (This image may be licensed either as rights managed or royalty free.)
RF - Lodgepole pines (Pinus contorta) covered in heavy early morning hoar frost. Yellowstone National Park, USA. January. Digitally stitched image
Red fox (Vulpes vulpes) walking through deep winter snow. Hayden Valley, Yellowstone, USA. January
Coyote (Canis latrans) standing in snow, dwarfed by frosty trees. Yellowstone National Park, USA, January
RF - Petrified trees and travetine cascade. Geothermal feature at Mammoth Hot Springs. Yellowstone National Park, USA. (This image may be licensed either as rights managed or royalty free.)
RF - Red fox (Vulpes vulpes) in deep snow. Yellowstone National Park, USA. January (This image may be licensed either as rights managed or royalty free.)
RF - North American river otters (Lutra canadiensis) on the frozen river edge. Yellowstone National Park, USA. January (This image may be licensed either as rights managed or royalty free.)
Coyote (Canis latrans) foraging in deep winter snow disturbed by grazing American bison (Bison bison) Hayden Valley, Yellowstone National Park, Wyoming, USA. January
North American river otters (Lutra canadiensis) probaly female with two juveniles, on the frozen river edge. Upper Yellowstone River, Hayden Valley, Yellowstone, USA. January
American Bison (Bison bison) frost covered male, standing by dead trees. Firehole River Valley. Yellowstone National Park, Wyoming, USA. January
RF - Herd of American Bison (Bison bison) around geo-thermal features. Firehole River Valley. Yellowstone National Park, Wyoming, USA. January 2008
Sun beam at sun rise over the Grand Canyon of the Yellowstone, Yelllowstone National Park, Wyoming, USA. January 2016
Grizzly bear (Ursus arctos horribilis) female with sub-adult cub rolling on back, in grassland. Yellowstone National Park, Wyoming, USA. June
RF - Red fox (Vulpes vulpes) foraging in snow covered valley. Hayden Valley, Yellowstone National Park, USA. February 2019. (This image may be licensed either as rights managed or royalty free.)
Steam rising from Grand Prismatic thermal pool in snow covered landscape. Midway Geyser Basin, Yellowstone National Park, USA. February 2019
Coyote (Canis latrans) standing near waters edge, in snow. Hayden Valley, Yellowstone National Park, USA. January 2019
Wolf (Canis lupus) lone male walking in shadow of the Teton Mountains. Snake River valley near Jackson Hole. Grand Teton National Park, Wyoming, USA. January
Coyote (Canis latrans) jumping in snow with its prey, Yellowstone National Park, Wyoming, USA, February
American elk (Cervus elaphus canadensis) stag sticking tongue out, Yellowstone National Park, Wyoming, USA
North American Bobcat (Lynx rufus) stalking along Madison River. Yellowstone National Park, Wyoming, USA. January
Pronghorn (Antilocapra americana) male, Yellowstone National Park, Wyoming, USA, June
American Buffalo or Bison (Bison bison) group with calves, Yellowstone National Park, Wyoming, USA, May
Bighorn sheep (Ovis canadensis) close up rams head shot, Yellowstone National Park, USA, February
Black bear (Ursus americanus) young cub portrait, Yellowstone National Park, Wyoming, USA
Black bear (Ursus americanus) cub, brown phase, looking out from behind tree trunk, Yellowstone National Park, Wyoming, USA, May
North American river otter (Lontra canadensis) playing in snow, Yellowstone National Park, Wyoming, USA, February
RF -Three Bison (Bison bison) walking through snow with herd feeding in background. Yellowstone National Park, Wyoming, USA. January
Coyote (Canis latrans) in snow, Yellowstone National Park, Wyoming, USA, February
Red fox (Vulpes vulpes) resting and sitting on snow, Yellowstone National Park, USA, February
American buffalo (Bison bison) with group of calves, Yellowstone National Park, Wyoming, USA. May
Bighorn Sheep (Ovis canadensis) Yellowstone National Park, Wyoming, USA. January
Bison (Bison bison) standing steam from geothermal springs, Yellowstone National Park, Wyoming, USA, February
