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Scarce Copper (Lycaena virgaureae) male in meadow on ox-eye daisy, Joutsa (formerly Leivonmaki), Finland, July
Arctic Fox (Alopex / Vulpes lagopus) at sunset, during moult from grey summer fur to winter white. Dovrefjell National Park, Norway, September
Marsh harrier (Circus aeruginosus) adult female circling over Oilseed rape crop, Norfolk, England, May. Digital composite
Keel billed toucan (Ramphastos sulfuratus), captive
Scenic view of a bull kelp forest (Nereocystis luetkeana) with sunlight shining through the fronds. Browning Pass, Port Hardy, Vancouver Island, British Columbia. Canada. North East Pacific Ocean
Lows cymbidium (Cymbidium lowianum) endemic tree orchid, Gaoligongshan NP, Yunnan province, China
Large white butterfly (Pieris brassicae), Vallee de l Eure (Eure Valley), Eure-et-Loir, France, August
Appennine Trumpet Gentian (Gentiana dinarica) in flower, on roadside below Gran Sasso, Appennines, Abruzzo, Italy, May
Clown frog (Dendropsophus leucophyllatus), Villa Carmen Biological Station, Peru
Nine-spotted moth (Amata phegea) feeding from a Mountain arnica flower (Arnica montana) Triglav National Park, Julian Alps, Slovenia. July
English oak (Quercus robur) acorn. Cambridgeshire, UK. September
Buff-tailed Bumblebee (Bombus terrestris) worker feeding on Ox-eye Daisy (Leucanthemum vulgare). Peak District National Park, Derbyshire, UK. July
Indian moon / Indian luna Moth (Actias selene) close up of defensive spines on caterpillar. Captive
Shelduck (Tadorna tadorna) female with ducklings, Cley, Norfolk, East Anglia, England, UK, July
Juvenile Peregrine falcon (Falco peregrinus) vocalising whilst perched in the Avon Gorge, Bristol, England, UK, June
RF- Pair of Rock beauties (Holacanthus tricolor) spawning at dusk. The larger male rubs against the flank of female during the spawning rise. Cayman Islands, British West Indies. Caribbean Sea
RF- Pair of Red Sea anemonefish (Amphiprion bicinctus) in Magnificent sea anemone (Heteractis magnifica), which has closed up in the late afternoon revealing its red skirt. St Johns Reef. Egypt
Butterfish (Pholis gunnellus) portrait. Gardur, south west Iceland. North Atlantic Ocean
Secretary blenny (Acanthemblemaria maria) making a home in a deserved worm tube of a great star coral (Montastraea cavernosa), East End, Grand Cayman, Cayman Islands, British West Indies
Male Chaffinch (Fringilla coelebs) perched on Gorse, Suffolk, UK, April
Male Dartford warbler (Sylvia undata) singing on gorse, Dorset, UK, April
Great white egret (Ardea alba) silhouetted at sunset, Lake Csaj, Pusztaszer, Hungary, February. Winner of the Portfolio category of the Terre Sauvage Nature Images Awards competition 2015
Hedge Parsley seed head (Torilis japonica) silhouetted at sunset, Norfolk, UK, November
Bullseye / Io moth (Automeris io) showing eye spot markings on wings during deimatic display to deter predators, originating from North and Central America, sequence 2 of 2. Captive
Orange cup coral (Tubastrea coccinea) Malpelo Island National Park, UNESCO World Heritage Site, Colombia, East Pacific Ocean
Brimstone moth (Opisthograptis luteolata) Banbridge, County Down, Northern Ireland, June
Aster flowers (Asteraceae sp) Vosges, France, May
Chinese moon moth (Actia sinensis subaurea) occurs in Northern China
Sunflowers (Helianthus annuus) in bloom, Norfolk, England, UK, August
Ruppells weaver (Ploceus galbula) male building nest, Oman, May
Common / Eurasian cranes (Grus grus) taking flight for roasting site, at sunset, silhouetted. Lake Hornborga, Sweden, April
Coral clam (Pedum spondyloideum) close up, Maldives, Indian Ocean
Galapagos land iguana (Conolophus subcristatus) skin and scales close-up. Isabela Island, Galapagos, Ecuador, June
Wood Warbler (Phylloscopus sibilatrix) singing from an oak tree, Atlantic Oakwoods of Sunart, Scotland, May
Lesser celandine (Ranunculus ficaria) close up of flowers, Broxwater, North Cornwall, UK, March
A tiny amphipod (Iphimedia obesa) living on Dead mans fingers soft coral (Alcyonium digitatum) Loch Carron, Ross and Cromarty, Scotland, UK, April
RF- Daffodils (Narcissus) in flower photographed using soft focus technique
Eyelash Palm-pitviper (Bothriechis / Bothrops schlegeli) coiled in strike pose with tongue extended. Costa Rica. Captive
Kelp gull (Larus dominicanus) on rocky shore with crashing waves at sunset, Bird Island, Lamberts Bay, Western Cape province, South Africa, September 2012
Female Downy Emerald dragonfly (Cordulia aenia) at rest on flowering Gorse (Ulex europaeus) Higher Hyde Heath, Dorset, UK. April
Striped triplefin (Helcogramma striata) on hard coral, Tubataha Reef, Philippines
Two Great egrets (Ardea alba) wading, silhouetted at dawn, Keoladeo National Park, Bharatpur, Rajasthan, India
Spinyhead blenny (Acanthemblemaria spinosa) in hard coral, Netherlands Antilles, Bonaire, Caribbean
Yellow flag iris (Iris pseudacorus), Bchelberg, Pfalz, Germany. May. Meetyourneighbours.net project
Brimstone butterfly (Gonepteryx rhamni) Male at rest on Primrose flower, West Sussex, UK April
American Bison (Bison bison) in habitat, Grand Teton National Park, Wyoming, USA, October
Great Tit (Parus major) perched on branch with yellow from autumn trees in the background. Southern Norway, October
Masai giraffes (Giraffa camelopardalis tippelskirchi), at sunrise, Masai-Mara Game Reserve, Kenya. March