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Red squirrel (Sciurus vulgaris) with autumn colours, Cairngorms National Park, Highlands, Scotland, UK. October
Flowering machair, South Uist, Outer Hebrides, Scotland, UK
Buttercup (Ranunculus acris) flowering in machair grassland, South Uist, Outer Hebrides, Scotland, UK, July
Silver birch (Betula pendula) woodland in early autumn, Craigellachie National Nature Reserve, Aviemore, Cairngorms National Park, Scotland, UK, October 2014
Willow grouse (Lagopus lagopus) Kiilopaa, Inari, Finland, January
Mirador de Morro Velosa (650 meters) Fuerteventura, Canary Islands. April 2013
Juvenile Banggai cardinalfish (Pterapogon kauderni) with sea anemone (Heteractis magnifica). Lembeh Strait, North Sulawesi, Indonesia
Black necked grebe (Podiceps nigricollis) pair at night, performing their courtship dance during the mating season, with street lights reflected in the water. The Netherlands.April 2014
Maple leaves backlit on lightbox, Broxwater, Cornwall, UK. November 2013
Great White Egret (Egretta alba) with water splashing from beak, Hungary January
Autumnal foliage of Fireweed (Epilobium angustifolium) Kluane National Park, Yukon Territories, Canada, September
African Blue tit (Cyanistes teneriffae) Morocco, February
Lioness (Panthera leo) and Thomsons gazelles (Eudorcas thomsonii) silhouetted at sunset, Masai-Mara game reserve, Kenya
Four Yellow-billed storks (Mycteria ibis) standing in water, Lake Nakuru, Nakuru National Park, Kenya
European Herring Gull (Larus argentatus) flock in flight at sunset Norway, May
Red lechwe (Kobus leche) silhouetted at sunrise, Okavango Delta, Botswana
Split level view of shallow coral reef in the Red Sea at sunset. Gordon Reef, Sinai, Egypt. Strait of Tiran, Gulf of Aqaba, Red Sea
King Penguins (Aptenodytes patagonicus) on beach at sunrise, South Georgia Island, March
Male Long tailed duck (Clangula hyemalis) on water, Batsfjord, Norway, March
Male King eider duck (Somateria spectabilis) on water, Batsfjord, Norway, March
Conifers reflected in water, Strbske Pleso, Tatra Mountains, Slovakia, June 2013
Rock formations silhouetted at sunset on the Pacfic coast of Olympic National Park, Washington State, USA
Cowslips (Primula veris) growing on organic farm. Norfolk, UK, April
Mallard (Anas platyrhynchos) walking on ice, Acadia National Park, Maine, USA, April
Moorland Clouded Yellow butterfly (Colias palaeno) male and female, Joutsa (formerly Leivonmaki), Finland, July
Whooper Swan (Cygnus cygnus) male and female facing in opposite directions, central Finland, April
Northern Chequered Skipper, (Carterocephalus silvicola), female on flower
Painted lady butterfly (Cynthia / Vanessa cardui) feeding on Fleabane flower, UK, August
Silver-studded blue butterfly (Plebejus argus) pair mating, resting on a Pyrimidal orchid (Anacamptis pyramidalis) flower, Gwithian Towans, Cornwall, UK, July
Harts tongue fern (Phyllitis scolopendrium) leaf unfurling, Cornwall, UK, May
Cart tracks in a flowering meadow, with Cork oaks (Quercus suber) in the background, Beja, Portugal, April
Shining honeycreeper (Cyanerpes lucidus) Costa Rica
Herd of Impala (Aepyceros melampus) silhouetted at sunset, Ngorongoro Conservation Area, Tanzania
California condors (Gymnnogyps californicus) interacting. Captive. Endangered species
Fissure eruption of Chico Volcano into 9km diameter caldera. Sierra Negra volcano, Isabela Island, Galapagos, Ecuador. October 2005
Candy stripe shrimp (Lebbeus grandimanus) on a yellow sponge. Browning Pass, Port Hardy, Vancouver Island, British Columbia. Canada. North East Pacific Ocean
Valdivia Ground Frog 1+Eupsophus vertebralis+2 Oncol Park, Chile, January
A portrait of a large male Dusky grouper (Epinephelus marginatus). Lavezzi Islands, between Sardinia and Corsica in the Mediterranean Sea
Scarlet Pimpernel (Anagallis foemina) blue form close-up of stamens, in a garden, Orvieto, Italy, May
Swallowtail (Papilio machaon) in flight, Finland, Europe
Flooded farmland with hay bales on Tadham Moor, Somerset Levels, England, December 2012
Sunrise behind Mount Mikeno, Virunga National Park, Democratic Republic of the Congo, August 2010
Beluga whale (Delphinapterus leucas) swimming under ice and exhaling air, Arctic circle Dive Center, White Sea, Karelia, northern Russia. Captive
Female goshawk (Accipiter gentilis) in flight, just after taking off from perch. Southern Norway, January
Common / Eurasian crane (Grus grus) calling at sunset, silhouetted. Lake Hornborga, Sweden, April
Kea (Nestor notabilis) portrait. The yellow at the top of the beak indicates that this is a jurvnile bird. Arthurs Pass National Park, South Island, New Zealand, December
Fallow deer (Dama dama) one deer stops and looks whilst the herd runs on, with sunlit leaves out of focus, Vastmanland, Sweden
Red Sea anemonefish (Amphiprion bicinctus) in sea-anemone, Red Sea
