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Arctic Fox (Alopex / Vulpes lagopus) yawning, during moult from grey summer fur to winter white. Dovrefjell National Park, Norway, September
Giant panda (Ailuropoda melanoleuca) cub yawning. Yuan Meng, first giant panda ever born in France, age 10 months, Captive at Beauval Zoo, Saint Aignan sur Cher, France
Clarks Grebe (Aechmophorus clarkii) with yawning chick riding on back, Bear River Refuge, Utah, July
Tasseled wobbegong shark (Eucrossorhinus dasypogon) yawning. Citrus Ridge, Yanggefo Island, Gam Island, Raja Ampat, West Papua, Indonesia
Eurasian lynx (Lynx lynx) yawning. Captive
Javan leopard (Panthera pardus melas) roaring, native to the Indonesian island of Java. Captive. Digital composite
Jaguar (Panthera onca) female yawning, native to Southern and Central America, captive
Golden Retriever dog pup, Oscar, 3 months, in play-bow, against white background
Large male Napoleon wrasse (Cheilinus undulatus) yawning. Ras Mohammed, Sinai, Egypt. Red Sea
Leopard (Panthera pardus) female yawning, Samburu game reserve, Kenya
Smooth coated Jack Russell Terrier, black, tan and white, female with two 8 week puppies
Frilled shark (Chlamydoselachus anguineus) specimen with mouth open, from Atlantic Ocean, at a depth of 729m
Wild goats (Capra hircus) resting on coastal promontory, one yawning, Great Orme, Llandudno, Gwynedd, North Wales, UK, August
Head portrait of Sumatran tiger (Panthera tigris sumatrae) with mouth wide open in a yawn, captive
Black grouper (Mycteroperca bonaci) yawning on a coral reef. Jardines de la Reina, Gardens of the Queen National Park, Cuba. Caribbean Sea
Red fox (Vulpes vulpes) fox waking up from sleep in den in deep snow, stretching and yawning, Kamchatka, Far east Russia, November
Clouded leopard (Neofelis nebulosa) resting in tree, Tripura state, India. Captive
Japanese Macaque (Macaca fuscata) male yawning, Jigokudani Japan, January
Whale shark (Rhincodon typus) mouth open filter feeding at the surface, Quintana Roo, Yucatan Peninsular, Mexico, Caribbean Sea
Secretary blenny (Acanthemblemari maria) yawns as it peers out from a hole in the reef. West Bay, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
Konik Wild Horse (Equus ferus caballus) yawning. The Netherlands, November
Spotted hyaena laughing {Crocuta crocuta} Kenya
Adult male Hippopotamuses (Hippopotamus amphibius) posturing in agressive yawn behaviour, Luangwa River, South Luangwa National Park, Zambia, November. Digital composite (two frames stitched together)
Red fox (Vulpes vulpes) vixen yawning in the morning light (captive). Lifton, Devon, UK, May
Arctic fox yawning {Vulpes lagopus} Norway captive
Two Lionesses (Panthera leo) in the rain, one shaking water off and the other yawning. Masai-Mara game reserve, Kenya
Adult male Hippopotamus (Hippopotamus amphibius) posturing in agressive yawn behaviour, Luangwa River, South Luangwa National Park, Zambia, November
Grey seal (Halichoerus grypus) pup yawning. Donna Nook, Lincolnshire, UK. November
RF - Puffin (Fratercula arctica) standing on rock, yawning, Runde, Norway. June. (This image may be licensed either as rights managed or royalty free.)
Warty frogfish (Antennarius maculatus) yawning in its hiding place amongst white sponges, Bitung, North Sulawesi, Indonesia, Molucca Sea
Male Cheetah (Acinonyx jubatus) stretching and yawning on a fallen tree trunk, Okavango Delta, Botswana, Africa
South Georgia pintail (Anas georgica georgica) yawning, Ocean Harbour, South Georgia Island
Southern elephant seal (Mirounga leonina) male, rests and yawns on the snow. Right Whale Bay, South Georgia Island
African lion (Panthera leo) with yawning cub, Masai Mara National Reserve, Kenya, January
European red fox (Vulpes vulpes crucigera) in deep snow in front of steep rocks. Gran Paradiso National Park, Italy. December Highly commended in the Portfolio category of the Terre Sauvage Nature
Tiger (Panthera tigris), yawning, Ranthambhore National Park, Rajasthan, India
Snowy owl (Bubo scandiacus) yawning or bill-stretching, Grays Harbor County, Washington, USA December
Three European pond turtles (Emys orbicularis) and a Balkan terrapin (Mauremys rivulata) on a rock surrounded by water, Butrint, Albania, June 2009 WWE OUTDOOR EXHIBITION
Feral domestic rabbit (Oryctolagus cuniculus) standing on hind legs on coast, Okunojima Island, also known as Rabbit Island, Hiroshima, Japan
Giant river otter (Pteronura brasiliensis) yawning at water surface, Yasuni National Park, Orellana, Ecuador, Endangered species
Tabby kitten, Picasso, 7 weeks, yawning
Cheetah (Acinonyx jubatus) male yawning, Kruger national park, South Africa
Cheetah (Acinonyx jubatus) cub aged 6 months yawning, Masai-Mara Game Reserve, Kenya. Vulnerable species
Blue Bearded Collie puppy, 3 months, yawning
Border Collie with one of her puppies, 5 weeks, yawning
Wild stallion (Equus caballus) lying down in the sagebrush and yawning. The foothills south of Reno, Nevada, USA, May
Alpine marmot (Marmota marmota) calling, Hohe Tauern National Park, Austria, July
