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Brown bear (Ursus arctos) and Grey wolf (Canis lupus) together in wetlands, Kuhmo, Finland, July
Pair of European grey wolves (Canis lupus) interacting, Tromso, Norway, captive, April
European grey wolf (Canis lupus) interacting with a European brown bear (Ursus arctos) Kuhmo, Finland, July 2009. WWE INDOOR EXHIBITION
Wolf (Canis lupus) pack running in deep snow, captive in enclosure of the Bavarian Forest National Park, Germany, February
Grey wolf howling, USA
Pack of European grey wolves standing in snow (Canis lupus) captive. Bayerischerwald National Park, Germany
Head portrait of a Grey Wolf (Canis lupus) captive
Two European grey wolves (Canis lupus) howling in winter landscape, captive. Bayerischerwald National Park, Germany
European grey wolf (Canis lupus) running in snow captive. Bayerischerwald National Park, Germany
European Wolf (Canis lupis) adult female in boreal birch forest in winter, Bardu, Norway, captive
European Grey Wolf male yawning {Canis lupus} captive, Transsylvania, Romania
Arctic wolf running in snow {Canis lupus arctos} captive USA
Grey wolves showing fur colour variation. (Canis lupus)
Portrait of a wild Carpathian Grey Wolf (Canis lupus lupus) in snow-bound woodland habitat. Bieszczady, Carpathian Mountains, Poland, December
Wild Grey wolf (Canis lupus) walking, Kuhmo, Finland, September
Portrait of European grey wolf (Canis lupus) in snow, Bayerisherwald National Park, Germany, Captive
Two European Grey wolves (Canis lupus) one howling, in snow, captive, Bayerisch National Park, Germany
Iberian wolf (Canis lupus signatus) captive, Spain
Arctic grey wolf {Canis lupus arctos} running through snow, USA. captive
European wolf (Canis lupis) in boreal birch forest in winter, Nord-Trondelag, Norway. Captive release
Grey wolf portrait with autumn leaves {Canis lupus} captive, USA
Grey wolves playing in snow {Canis lupus} captive, USA
Grey wolf head portrait {Canis lupus} US Captive
Grey wolf white Arctic form wild (Canis lupus) pup begging for food from mother. Ellesmere
Grey wolf (arctic form) with pup near den, Ellesmere Island, Canada
Grey wolf {Canis lupus} head profile, captive, Montana, USA
Grey Wolf (Canis lupus) head portrait of male, in snow, Captive
Portrait of Iberian wolf {Canis lupus sygnatus} snarling, captive, Lobo Park, Antequera, Malaga, Spain
European grey wolf (Canis lupis) in snow-laden boreal birch forest, Nord-Trondelag, Norway, captive
Arctic Grey wolf in snow {Canis lupus} captive, Idaho, USA
Grey wolf portrait {Canis lupus} captive, Minnesota, USA
European / Grey wolf (Canis lupus) at sunset, Kuhmo, Finland, July 2009. WWE INDOOR EXHIBITION
Wild Eurasian brown bear (Ursus arctos) and a European Grey wolf (Canis lupus) at waters edge, Kuhmo, Finland, July 2008
Wild European Grey wolf (Canis lupus) walking, Kuhmo, Finland, July 2008
Wild European Grey wolf (Canis lupus) silhoutted in mist, Kuhmo, Finland, July 2008
Wild European Grey wolf (Canis lupus) Kuhmo, Finland, September 2008
Wolves (Canis lupus) sitting on a hillside in snow. Yellowstone, USA, February
Eurasian / European / Forest Wolf (Canis lupus lupus) head portrait resting on ground. Mongolia, August
Grey wolf (Canis lupus) walking through water, Yellowstone National Park, USA, North America, May
Adult Arctic wolf (Canis lupus) Ellesmere Island, Nunavut, Canada, June 2008. Taken on location for BBC series, Frozen Planet, Summer
Arctic wolf (Canis lupus) on tundra, Ellesmere Island, Nunavut, Canada, June 2008. Taken on location for BBC series, Frozen Planet, Summer
Grey wolf (Canis lupus) close up portrait, captive
Grey wolf (Canis lupus) close up, captive
Portrait of Grey Wolf (Canis lupus). Yellowstone National Park, Wyoming, USA, October
Grey wolf (Canis lupus) head portrait, viewed through autumn colours, captive
Grey Wolf (Canis lupus) head portrait of male, lying in snow, Captive
Wild Grey wolf (Canis lupus) Kuhmo, Finland, September
Two Grey wolf (Canis lupus) cubs standing together in green vegetation, captive
