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Least Chipmunk (Tamius minimus) feeding on the seeds of a thistle bush during a snowstorm, Grand Teton NP, Wyoming, USA, October
White-winged Dove (Zenaida asiatica), adult perched on icy branch of Yaupon Holly (Ilex vomitoria), Hill Country, Texas, USA. February
Western Scrub-Jay (Aphelocoma californica), adult calling on icy branch of Possum Haw Holly (Ilex decidua) with berries, Hill Country, Texas, USA. February
Male Eastern bluebird (Sialia sialis) feeding on winterberry (Ilex sp) fruit, New York, USA, January
Bald Eagle (Haliaeetus leucocephalus) adult, winter, Kenai Peninsula, Alaska, USA
RF- Capercaillie (Tetrao urogallus) male displaying in heavy snowfall, Cairngorms National Park, Scotland. February
Small stream flowing through snow covered landscape, Forollhogna National Park, Norway, September 2008
European bison (Bison bonasus) herd walking through an agricultural field, Bialowieza NP, Poland, February 2009
Two Muskox (Ovibos moschatus) head to head, Dovrefjell National Park, Norway, February 2009. WWE BOOK
Muskox (Ovibos moschatus) feeding, Dovrefjell National Park, Norway, February 2009
Two Muskox (Ovibos moschatus) on the horizon, Dovrefjell National Park, Norway, February 2009
Five Muskox (Ovibos moschatus) lying down, in mist, Dovrefjell National Park, Norway, February 2009
Two Muskox (Ovibos moschatus) in snow, Dovrefjell National Park, Norway, February 2009
Muskox (Ovibos moschatus) close-up, Dovrefjell National Park, Norway, February 2009. WWE INDOOR EXHIBITION
Two Muskox (Ovibos moschatus) face to face, Dovrefjell National Park, Norway, February 2009
Rear view of Muskox (Ovibos moschatus) behind snow bank, Dovrefjell National Park, Norway, February 2009
Photographers camp, Dovrefjell National Park, Norway, February 2009
Rear view of Muskox (Ovibos moschatus) grazing with snow on its back, Dovrefjell National Park, Norway, February 2009
Rear view of Muskox (Ovibos moschatus) lying in snow, Dovrefjell National Park, Norway, February 2009
Low clouds over snow covered hills, Dovrefjell National Park, Norway, February 2009
Willow grouse (Lagopus lagopus) in snow, Dovrefjell National Park, Norway, February 2009
Mountain in Dovrefjell National Park, Norway, February 2009
Willow grouse (Lagopus lagopus) on rock in snow, Dovrefjell National Park, Norway, February 2009
Mountain hare (Lepus timidus) footprints in snow, Creag Meagaidh National Nature Reserve, Scotland, UK, December 2012
Fieldfare (Turdus pilaris) perched on branch of a Rowan tree (Sorbus aucuparia), with berries, Clwyd, Wales, UK, November
Flock of Dark-bellied brent geese (Branta bernicla) flying over sea wall, South Swale, Kent, England, UK
Forest of scots pine after heavy snowfall, Cairngorms National Park, Scotland, March 2012. Did you know? Like many orchids
Snow-covered Beech (Fagus sylvatica) woodland. West Woods, Compton Abbas, Dorset, England, UK, December
Beech (Fagus sylvatica) woodland in hoar frost and winter mist. West Woods, Compton Abbas, Dorset, England, UK, December
Short-eared owl (Asio flammeus) flying with dead vole held in its beak, Worlaby Carr, Lincolnshire, England, UK, December
Barnacle geese (Branta leucopsis) feeding on grazing marshes, with one bird wing stretching, Caerlaverock WWT, Dumfries and Galloway, Scotland, UK, December
Scots pine (Pinus sylvestris) woodland in winter, Abernethy NNR, Cairngorms NP, Scotland, UK, December 2011
European river otter (Lutra lutra) footprints crossing frozen Loch Lurgainn at dawn, with Sgorr Tuath & Sgorr Deas (Beinn an Eoin) behind, Coigach, Wester Ross, Scotland, UK, December 2010
Cul Beag in winter, Coigach, Wester Ross, Scotland, UK, December 2010
Female Blackbird (Turdus merula) perched in crab apple tree in winter, Scotland, UK, December 2010. Did you know? The oldest ringed blackbird recovered was over 20 years old!
Male Blackbird (Turdus merula) perched on garden seat in winter, with feathers ruffled to insulate against cold, Scotland, UK, December 2010
Mountain hare (Lepus timidus) in winter coat sitting in the snow, Scotland, UK, February
Mountain hare (Lepus timidus) scratching for food in the snow, Scotland, UK, February
Red deer stag (Cervus elaphus) walking on open moorland in snow, Cairngorms NP, Scotland, UK, December
Group of Mute swans (Cygnus olor) on a partially frozen loch, Loch Laggan, Creag Meagaidh NNR, Scotland, UK, December 2010
Windswept Silver birch trees (Betula pendula) silhouetted against sky, Cairngorms National Park, Scotland, UK, November
Fish-eye image of Scots pine trees (Pinus sylvestris) in pine forest, Abernethy forest, Highland, Scotland, UK, February
Dead Scots pine tree (Pinus sylvestris) laden with snow in winter, Rothiemurchus Forest, Cairngorms NP, Highland, Scotland, UK, February 2010
Red squirrel (Sciurus vulgaris) on pine stump in snow, Scotland, UK, December. Did you know? Red squirrels can locate their buried food supplies in over a foot of snow!
Rock ptarmigan (Lagopus mutus) male hidden by rocks, eye visible, winter plumage, Cairngorm Mountains, Cairngorms NP, Highland, Scotland, UK, February
Mist swirling around Coire an Lochain in winter, Cairngorm Mountains, Cairngorms NP, Highland, Scotland, UK, February 2011
Winter sunset over the Somerset Levels from Walton Hill on a misty evening with snow on the ground. View includes Butleigh Moor, Somerton Moor, Street Moor, Low Ham Moor, Walton Moor, Pitney Moor
Manx Loaghtan Sheep (Ovis aries) used for grazing on unimproved grassland on Minsmere RSPB Reserve, habitat for breeding Stone Curlews, Suffolk Sandlings, Suffolk, UK, February 2011