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Porcelain toadstools (Oudemansiella mucida) New Forest, Hampshire, UK, October
Oak woodland with mosses and running water, near Dolgellau, Snowdonia, North Wales, UK October
Honey fungus (Armillaria borealis) growing on dead birch, Sherwood Forest, Nottinghamshire, UK October
Lime tree (Tilia sp) avenue in autumn, Hampstead Heath, London, England, UK. November
Chamois (Rupicapra rupicapra) Ballons des Vosges Regional Natural Park, Vosges Mountains, France, October
Pine trees in their landscape emerging from the clouds and mist, Ballons des Vosges Regional Natural Park, Vosges, France, October
Eurasian red squirrel (Sciurus vulgaris) reflected in water, Cairngorms National Park, Scotland, United Kingdom, October
View of Le Mont Pourri, with European larch (Larix decidua) forest, Le Monal Vanoise National Park, Savoie, France, October 2015
View up towards Beech (Fagus sylvatica) woodland canopy. Derbyshire, UK. October
Money spider (Linyphiidae) in its sheet web eating a beetle, backlit at sunset. Thursley Common National Nature Reserve, Surrey, UK. October
Small stream in forest, Vosges mountains, France, November 2014
Lapwing (Vanellus vanellus) Titchwell RSPB Reserve, Norfolk, England, UK. November
Autumnal forest, Vosges forest, France, November 2014
Autumn mist in Dee Valley (Dyffryn Dyfrdwy) near Llangollen, Denbighshire, Wales, UK, November 2013
Flock of Pink footed geese (Anser brachyrhynchus) in flight at dusk, Norfolk, England, UK, October
Autumnal Beech (Fagus) trees, Savernake Forest, Wiltshire, UK, November 2012
Red squirrel (Sciurus vulgaris) foraging in forest, climbing down tree trunk, Cairngorms National Park, Scotland, October
Wild cat (Felis silvestris) climbing tree in autumn, Black Forest, Baden-Wurttemberg, Germany. October
Landscape of the Verzasca River in autumn, Canton Tessin, Switzerland, November 2011. Taken for the Freshwater Project
Toadstoosl on tree stump in forest, Vosges, France, November 2013
Misty Beech (Fagus sylvatica) forest in autumn, Vosges mountains, France, November 2013
Fly Agaric (Amanita muscaria), Peak District National Park, Derbyshire, UK. October
Sheathed Woodtuft fungi (Kuehneromyces mutabilis) Surrey, England, UK, November
RF- Fallow deer (Dama dama) doe at sunrise during rut, Deer Park, Holkham, Norfolk, UK, October
Captive reared juvenile White stork (Ciconia ciconia) in flight over the Knepp Estate soon after release, Sussex, UK, August 2019
Wild cat (Felis sylvestris) portrait, part of captive breeding project at Alladale Estate, Scotland, UK, October
Purple vipers bugloss (Echium plantaineum) growing in fields of Corn Marigold (Chrysanthemum)Purple vipers bugloss (Echium plantaineum) growing in fields of Corn Marigold (Chrysanthemum segetum) Project Neptune National Trust, Cornwall, England, UK, June
Lesser horseshoe bats (Rhinolophus hipposideros) in magnesium mine, Shropshire, England, UK, April
Sequence of strips of cultivated Machair with early shoots of Black oats (Avena strigosa) pushing through the sand. North Uist, Scotland, UK, June
Corncrake (Crex crex) walking at dawn on Machair, North Uist, Scotland, UK, June
Two Little terns (Sterna albifrons) at the nest amongst Black oats (Avena strigosa) growing on shell rich sands Machair, North Uist, Scotland, UK, June
Little tern (Sterna albifrons) at the nest amongst Black oats (Avena strigosa) growing on shell rich sands Machair, North Uist, Scotland, UK, June
Little tern (Sterna albifrons) with newly hatched chick, Gronant dunes, Denbighshire, Wales, UK, June
Little tern (Sterna albifrons) nest with three eggs in shingle ridge. County Wicklow, Ireland, June
Corn Marigold (Chrysanthemum segetum) and Scentless mayweed (Tripleurospermum inodorum) machair habitat, North Uist, Scotland, UK, July
Red Deer (Cervus elaphus) portrait of head of stag and antlers. Alladale Estate, Scotland, UK, October
Blue tit (Cyanistes caeruleus) bringing Crab spider (Misumena vatia) prey to nest, Monmouthshire Wales, UK, May
Painted nomad bee (Nomada fucata) male bee with characteristic green eyes, sleeping with jaws locked around grass stem, Monmouthshire, Wales, UK. June
Smeathmans furrow bee (Lasioglossum smeathmanellum), feeding on wild alpine strawberry (Fragaria vesca), Monmouthshire, Wales, UK. December
Partially wooded uplands with Five Sisters of Kintail in background, Northwest Highlands, Scotland, UK.September
Purple loosestrife (Lythrum salicaria) flower spikes in evening light, Scotland, UK, September
Potrait of a Cuckoo wrasse (Labrus bimaculatus) Loch Linnhe, west coast of Scotland, UK, September
Hermit crab (Pagurus bernhardus) on yellow sponge, Loch Etive, west coast of Scotland, UK, September
Collared earth star (Geastrum triplex), two on woodland floor. South Yorkshire, England, UK. September
Chinese Water Deer (Hydropotes inermis) Norfolk UK September
Small copper butterfly (Lycaena phlaeas) standing on an Aster flower in a meadow, Bath and northeast Somerset, UK, September
Seaweeds including Thongweed / Sea thong (Himanthalia elongata), Tangleweed kelp (Laminaria digitata), Toothed wrack (Fucus serratus) and Dulse (Palmaria palmata)
Common Toad (Bufo bufo) juvenile toadlet walking over moss in birchwood in late summer, Inverness-shire, Scotland, September