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Lesser spotted fritillary (Melitaea trivia) adult at rest on yellow flower, Bulgaria
European robin (Erithacus rubecula) adult with autumn leaves in background, Punkaharju, Etela-Savo / Southern Savonia, Ita-Suomi / Eastern Finland, Finland. September
Male impalas (Aepyceros melampus) standing in heavy rain, Masai-Mara Game Reserve, Kenya. February
Black Grouse (Tetrao tetrix) male displaying with breath condensing in cold air, Kuusamo, Finland April
African Blue tit (Cyanistes teneriffae) Morocco, February
Portrait of male Waterbuck (Kobus ellipsiprymnus) in the rain, Masai-Mara game reserve, Kenya
Four Yellow-billed storks (Mycteria ibis) standing in water, Lake Nakuru, Nakuru National Park, Kenya
Three Short-finned pilot whales (Globicephala macrorhynchus) close to the surface against the light, Costa Rica. Pacific ocean
Hooded crow (Corvus coronix) and Eurasian Magpie (Pica pica) waiting to scavenging White tailed eagles (Haliaeetus albicilla) food, Pusztazer, Hungary
Small pearl bordered fritillary butterfly (Boloria selene) resting on grass, backlit underwing, Marsland Mouth, North Devon, UK, June
Small Yellow Underwing moth (Panemeria tenebrata) on stitchwort (Stellaria) South Karelia, southern Finland, June
Moth (Zygaena osterodensis) feeding on flower, Viscos, Pyrenees National Park, France, July
Painted lady butterfly (Cynthia / Vanessa cardui) feeding on Fleabane flower, UK, August
Bulldog puppy with chin on paws, against white background
Harvest mice (Micromys minutus), captive, UK, June
Herd of Impala (Aepyceros melampus) silhouetted at sunset, Ngorongoro Conservation Area, Tanzania
tail detail of Sperm Whale (Physeter macrocephalus) Pico Island, Azores, Portugal, Atlantic Ocean
Spectacled caiman (Caiman crocodilus) resting in shrinking pool, Pouso Alegre lodge. Pantanal, Brazil. Highly commended in the Other Animals Category of the GDT competition 2013
Valdivia Ground Frog 1+Eupsophus vertebralis+2 Oncol Park, Chile, January
Vanzolinis Spiny-chest Frog 1+Alsodes vanzolinii+2 on Eucalyptus leaves, endemic to the Nahuelbuta mountain range, Chile, December. Critically Endangered species
Komodo dragons (Varanus komodoensis) in forest clearing, Komodo Island, Indonesia
Gaoligongshan Tree Frog 1+Polypedates / Rhacophorus gongshanensis+2 in fern 1+Cibotium barometz+2 Gaoligongshan Mountains, Yunnan, China
Rufous elephant shrews (Elephantulus rufescens) Botswana
Hanuman / Northern Plains Grey Langur (Semnopithecus entellus) youngster sitting on the trunk of a Banyan tree, Ranthambore National Park, Rajasthan, India, June
Mountain Hare (Lepus timidus) feeding. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, March
Harlequin duck (Histrionicus histrionicus) male on a large boulder in the fast flowing and turbulent waters of the Laxa River, Iceland, May. Bookplate from Danny Greens The Long Journey North
European Robin (Erithacus rubecula) on snow covered birch tree branch, Vosges, France, January
Atlantic puffins (Fratercula arctica) at dusk, Fair Isle, Shetland, July. Bookplate from Danny Greens The Long Journey North
Puffin (Fratercula arctica) stretching wings, late evening light. Shetland Islands, Scotland, July
Red Sea anemonefish (Amphiprion bicinctus) in sea-anemone, Red Sea
Glass Frog (Cochranella mache) portrait, Ecuador, Endangered species
Aquatic Warbler (Acrocephalus paludicola). Zvanets fen mire, Belarus, Europe. Vulnerable Species
Lesser Spotted Woodpecker (Dendrocopus minor) on bullrush, Uto Finland October
Wolves (Canis lupus) sitting on a hillside in snow. Yellowstone, USA, February
Puffins (Fratercula arctica) among coastal thrift. Saltee Islands, Ireland, UK, July
RF - Common darter dragonfly (Sympetrum striolatum) resting on reed, silhouetted against the moon, Devon, UK. August
Hanuman langur / Northern Plains Grey Langur (Semnopithecus entellus) drinking from a water tap at Ranthambore fort. Ranthambore National Park, Rajasthan, India
Owlfly sulphur (Libelloides coccajus) on flower, near Gavarnie, Pyrenees National Park, France, June
Herd of Camargue bulls, Camargue, France, May
Golden Retriever, male in straw
Willow grouse (Lagopus lagopus) in winter plumage, Utsjoki, Finland, April
Male Long-tailed duck (Clangula hyemalis), Batsfjord, Norway, March
Leopard (Panthera pardus) back of head, Samburu National Reserve, Kenya
Kamchatka Brown Bear (Ursus arctos beringianus) stands on its back legs to more easily determine the source of an unfamiliar sound
Robin (Erithacus rubecula) in snow, Titchwell, Norfolk, January. Image digitally manipulated to add more snow
Dalmatian Pelicans (Pelecanus crispus) portrait. Lake Kerkini, Greece, March 2012
Red-eyed Treefrog (Agalychnis callidryas) on leaf, captive, from South America
Puffin (Fratercula arctica) at a breeding colony. Fowlsheugh, Scotland, June