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Ural Owl (Stix uralensis) resting in snowy tree, Kuusamo Finland February
Orcas / killer whales (Orcinus orca) swimming in open water, Three Kings Islands, New Zealand. Pacific Ocean
RF- Three baby lop rabbits
Willow Grouse (Lagopus lagopus) male perched on snow, Finland, March. Bookplate from Danny Greens The Long Journey North
RF - Red Squirrel (Sciurus vulgaris) on log in snow. Scotland, UK. December
Japanese Macaques (Macaca fuscata) in hot springs, Jigokudani, Nagano Prefecture, Honshu, Japan
The River Thames and Houses of Parliament at Westminster at sunset with gull, UK, November 2005
Hoatzins (Opisthocomus hoazin) perched in tropical rainforest, Tambopata Reserve, Peru, South America
Ginger and white kitten looking at camera
Golden-mantled tamarin (Saguinus tripartitus) captive
UK Ladybird species, native and invasive, from left to right: Seven-Spot (Coccinella 7-punctata), Harlequin (Harmonia axyridis), Harlequin morph, Two-Spot (Adalia bipunctata)
Golden Retriever x Poodle F1b Goldendoodle puppy on paw print background
Flock of White ibis (Eudocimus albus) in breeding plumage, at rookery on water sFlock of White ibis (Eudocimus albus) in breeding plumage, at rookery on waters edge of the mangrove-covered island, Tampa Bay, Florida, USA, March
Oceanic whitetip shark (Carcharhinus longimanus) accompanied by pilotfish (Naucrates ductor) as it cruises beneath the surface of the Red Sea in the late afternoon, close to Little Brother Island
Working English Springer Spaniel puppy, 6 weeks, in play bow
Black and white Border Collie puppy, sitting and looking up
Skewbald Horse in ranch, Martinsdale, Montana, USA
European common frog (Rana temporaria) on rock by waterfall, near Lac des Gloriettes dam, Pyrenees National Park, France, June
Black veined white butterfly (Aporia crataegi) on grass, Gavarnie-Gedre, Pyrenees National Park, France, June
Tree sparrows (Passer montanus) perching on a branch in the rain. Perthshire, Scotland, December
School of large Atlantic bluefin tuna (Thunnus thynnus) captive in growing pen. Depsite looking much smaller in the photo, these fish were 1.5m (5ft) long. Malta. Mediterranean Sea
The vertical reef wall at Shark Reef, Ras Mohammed, with Scalefin anthias (Pseudanthias squamipinnis) and soft corals (Dendronephthya spp.) Yellowsaddle goatfish (Parupeneus cyclostomus)
Marbled White butterfly (Melanargia galathea) resting on grass, Dunsdon Nature Reserve, Devon, UK. July
RF - School of large Atlantic bluefin tuna (Thunnus thynnus) captive in a growing pen. Depsite looking much smaller in the photo, these fish were 1.5m (5ft) long. Malta. Mediterranean Sea
RF - Eyelash Pit Viper (Bothriechis schlegelii) Costa Rica
Banded demoiselle (Calopteryx splendens) group of four on plant stems, River Leijgraaf, Nijmegen, the Netherlands, August 2013
Waterfowl at dawn with mist, including Great blue heron (Ardea herodias herodias), American avocet (Recurvirostra americana), Northern shoveler (Anas clypeata), Xochimilco wetlands, Mexico City
Kingfisher (Alcedo atthis) perched on branch, Black Forest, Baden-Wurttemberg, Germany
Pallas's cat (Otocolobus manul) kitten waiting at den for mother to return, Mongolia. June. 2017. Filmed for BBC series Big Cats
Humpback whale (Megaptera novaeangliae australis) female, stationary with calf heading to surface for breath of air. Vava'u, Tonga, South Pacific
RF- Fallow deer (Dama dama) doe at sunrise during rut, Deer Park, Holkham, Norfolk, UK, October
Female Snowy owl (Nyctea scandiaca) backlit surrounded by blowing snow, near Georgian Bay, Ontario, Canada
A Kamchatka Brown Bear (Ursus arctos beringianus) standing in breaking waves. Pacific coast of Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, July
African lion (Panthera leo), resting on a rock, surrounded by flies, Masai Mara Game Reserve, Kenya. November
RF- Head portrait of Giant panda (Ailuropoda melanoleuca) covered in snow, captive born in 2000
Wild rare Losino stallion standing in snow, Losa Valley, Burgos, Spain. January 2014
Whooper Swan (Cygnus cygnus) standing in snowfall, Hokkaido, Japan, February
Yellowhammer (Emberiza citinella) in winter hedgerow, Norfolk, England, UK, December
RF - Reticulated Glass Frog (Hyalinobatrachium valerioi) backlit showing highly translucent body. Osa Peninsula, Costa Rica
Western sandpipers (Calidris mauri) roosting on barrier island tidal flats. Terrebonne Parish, Louisiana, USA. October
RF - Portrait of domestic German shorthaired pointer pup in autumn. Pomfret, Connecticut, USA. October
Great white shark (Carcharodon carcharias) Guadalupe Island or Isla Guadalupe, Pacific Ocean, Mexico, September. Vulnerable species
Sheep (Ovis aries) resting on the Quiraing mountains with low cloud cover, Isle of Skye, Inner Hebrides, Scotland, UK. July
European hornet (Vespa crabro) worker in flight, Surrey, England, August
Coalition of male Lions (Panthera leo) in grassland before storm, Masai-Mara game reserve, Kenya
Sea otter (Enhydra lutris) floating on its back at surface among kelp, Alaska, USA. Gulf of Alaska, Pacific ocean
Marine iguana (Amblyrhynchus cristatus) swimming back to the surface, Galapagos. Pacific ocean