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Panther chameleon (Furcifer pardalis) sitting on branch with black background, Nosy Faly, Madagascar
Nudibranch (Janolus cristatus) Vela Luka, Korcula Island, Croatia, Adriatic Sea, Mediterranean
RF - Composite image of tropical nudibranchs on a black background showing variety and abundance of nudibranch species, Indo-Pacific
RF - Mantis shrimp (Odontodactylus scyllarus) on walk about on coral reef. Puerto Galera, Philippines
Nudibranch (Phyllidia marindica), Lembeh Strait, North Sulawesi, Indonesia. December
Cornflowers (Centaurea cyanus) growing on fallow fields near Orvieto, Umbria, Italy, June. Taken with fisheye lens
Two Red-footed booby (Sula sula) perched on branches, Genovesa Island, Galapagos Islands, Ecuador
Blue tropical reef fish composite image on black background, Blue triggerfish (Pseudobalistes fuscus), Bicolour angelfish (Centropyge bicolor), Elongate surgeonfish (Acanthurus mata)
Yellow tropical reef fish composite image on black background, Falcula butterflyfish (Chaetodon falcula), Yellow boxfish (Ostracion cubicus), Golden butterflyfish (Chaetodon semilarvatus)
Ultramel Okeetee corn snake, with recently laid eggs, an interspecies hybrid between a Corn snake, (Pantherophis guttatus), and a Grey rat snake (Pantherophis spiloides)
Golden mantella frog (Mantella aurantiaca) with freshly laid eggs on wet mossy ground
Desert rose (Adenium obesum) tree in flower, Bogoria Game Reserve, Kenya
RF- Glasswing butterfly (Greta oto), Costa Rica
Flock of White ibis (Eudocimus albus) in breeding plumage, at rookery on water sFlock of White ibis (Eudocimus albus) in breeding plumage, at rookery on waters edge of the mangrove-covered island, Tampa Bay, Florida, USA, March
Strawberry poison-dart frog, (Oophaga pumilio / Dendrobates pumilio), Costa Rica
Common tern (Sterna hirundo) in flight over water, Hampstead Heath, London, UK, August
RF - A vibrant Red Sea coral reef scene, with orange female Scalefin anthias fish (Pseudanthias squamipinnis) teeming over Fire coral (Millepora dichotoma)
Orange Tip Butterfly (Anthocharis cardamines) male resting on Bluebell (Endymion / Hyacinthoides nonscriptus). Peak DIstrict National Park, Derbyshire, UK. April
Spring Crocus (Crocus vernus) in flower in snow, Campo Imperatore, Gran Sasso, Appennines, Abruzzo, Italy, May
Toco Toucan (Ramphastos toco) head and beak profile portrait, Brazil
RF- Giraffe silhouette at sunset. Namibia, Etosha National Park
American goldfinch (Carduelis tristis) male in breeding plumage, perched in Eastern redbud flowers in spring, New York, USA May
RF - Sunflowers (Helianthus annuus) in bloom on Norfolk, England, UK. August
Forest reindeer, (Rangifer tarandus fennicus) in velvet, Viiksimo, Kuhmo region. Finland, July. Rare subspecies which were nearly extinct in the 19th Century
The Northern Lights / Aurora Borealis over a river, Vik, Iceland. February 2014
A split level image of Southern stingray (Hypanus americanus) swimming over a sand bar. Grand Cayman, Cayman Islands. Caribbean Sea. Digital composite
Mountain pansy (Viola lutea) flowers, Strathspey, Cairngorms National Park, Scotland, June
Borage (Borago officinalis) flowers, Menorca, Balearic Islands, Spain, Europe
Common poppies (Papaver rhoeas) backlit in evening light, Polly / Porth Joke, Pentire, Newquay, Cornwall, UK, July
Corn Marigold (Chrysanthemum segetum) and Cornflowers (Centaurea) in flower, July, England, UK
RF - Eyelash Pit Viper (Bothriechis schlegelii) Costa Rica
RF - European goldfinch (Carduelis carduelis) perched in field of Sunflowers (Helianthus annuus), Cadiz, Andalusia, Spain. July. (This image may be licensed either as rights managed or royalty free.)
Yellow wagtail (Motacilla flava) perched among flowering Rapeseed (Brassica napus), Scotland, UK. May
Panther chameleon (Furcifer pardalis) coiled tail close up on black background, Ambilobe, Madagascar
RF - Red-and-Green Macaws or Green-winged Macaws (Ara chloropterus) in flight over forest canopy. Buraco das Araras (Sinkhole of the Macaws), Jardim, Mato Grosso do Sul, Brazil. September
Arnica (Arnica montana) in flower on mountainside, Sibillini, Umbria, Italy. May
Blue / Rippled triggerfish (Pseudobalistes fuscus), adult and three juveniles, composite image on black background, Lembeh Strait, North Sulawesi, Indonesia, Indo-Pacific
Pfeffers flamboyant cuttlefish (Metasepia pfefferi) composite image on black background showing colour variations, Puerto Galera, Philippines, Indo-Pacific
Helmet jellyfish (Periphylla periphylla) drifting in the deep sea, Trondheimsfjord, Norway, Atlantic Ocean
Tail-spot blenny (Ecsenius stigmatura) peering out from amongst the coral, Triton Bay, West Papua, Indonesia, Pacific Ocean
Hairy frogfish / Striated frogfish (Antennarius striatus), portrait, Raja Ampat, West Papua, Indonesia, Pacific Ocean
Botany Bay diamond weevil (Chrysolopus spectabilis) close up, Toowoomba, Queensland, Australia
Crucifix toad / Holy cross frog (Notaden bennetti) sitting in shallow water after heavy summer rain, Westmar, Queensland, Australia
Weedy seadragon (Phyllopteryx taeniolatus) swimming over seaweeds. Sydney, New South Wales, Australia. Pacific Ocean
Snakelocks anemone (Anemonia viridis) growing on a stalk of Eelgrass (Zostera marina) on the seabed, Isle of Coll, Inner Hebrides, Scotland, UK, Atlantic Ocean. August
Large flower of the parasitic plant Rafflesia pricei, growing in rainforest, Mount Kinabalu, Borneo, May, 2013. Filmed for the BBC series Wonders of The Monsoon
European bee-eater (Merops apiaster) two perched on branch, Lower Silesia, Poland